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Idaho bladderpod

arctic bladderpod

Habit Perennials; caudex simple, (often enlarged by persistent leaf bases); densely pubescent, trichomes (sessile or short-stalked), rays furcate or bifurcate, (nearly smooth to finely tuberculate). Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth).
Stems

simple from base, decumbent, (occasionally few-branched), 0.5–1.5(–2) dm.

simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm.

Basal leaves

blade elliptic to broadly obovate, triangular, rhombic, or round, 1.5–3(–4) cm, margins often sinuate or shallowly lobed.

(usually ± rosulate);

blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire.

Cauline leaves

(sessile or shortly petiolate);

blade elliptic to oblanceolate to obovate, 0.5–1.5 cm, (base narrowed to petiole), margins entire.

(sessile or shortly petiolate);

blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire.

Racemes

compact to loose.

loose.

Flowers

sepals oblong to broadly elliptic, 4–7.5 mm, (lateral pair saccate or not);

petals spatulate, 7.5–10 mm.

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw).

Fruiting pedicels

(ascending or divaricate-spreading, straight to loosely sigmoid or curved), 4–10 mm.

(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout).

Fruits

(sessile or substipitate), elliptic, suborbicular, or oblong, strongly compressed (angustiseptate), 5–9 mm, (rounded to sharply keeled on 1 side, edges ± keeled);

valves: (margins covering replum edges or not), usually pubescent throughout or, rarely, glabrous inside;

ovules (4–)8–14(–16) per ovary;

style 2–4.5(–5) mm.

subglobose to ellipsoid, uncompressed, 4–6(–9) mm;

valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile;

replum as wide as or wider than fruit;

ovules (8–)10–14(–16) per ovary;

style 1–2.5(–4) mm.

Seeds

slightly flattened.

plump.

2n

= 60.

Physaria carinata

Physaria arctica

Phenology Flowering May–Aug.
Habitat Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance
Elevation 0-1800 m (0-5900 ft)
Distribution
from FNA
ID; MT; WY
[BONAP county map]
from FNA
AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia)
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Differences in fruit morphology become blurred and the three subspecies are often indistinguishable where their ranges meet near the intersection of Idaho, Montana, and Wyoming.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits elliptic, not keeled, valve margins (thin, rounded), not covering replum edges.
subsp. paysonii
1. Fruits elliptic, suborbicular, or broadly oblong, keeled on one side, valve margins covering replum edges
→ 2
2. Fruits elliptic, bases narrow-rounded to acute, apices ± acute.
subsp. carinata
2. Fruits suborbicular to elliptic or broadly oblong, bases and apices rounded.
subsp. pulchella
Source FNA vol. 7, p. 629. FNA vol. 7, p. 626.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. carinata subsp. carinata, P. carinata subsp. paysonii, P. carinata subsp. pulchella
Synonyms Lesquerella carinata Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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