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Idaho bladderpod

threadleaf bladderpod

Habit Perennials; caudex simple, (often enlarged by persistent leaf bases); densely pubescent, trichomes (sessile or short-stalked), rays furcate or bifurcate, (nearly smooth to finely tuberculate). Annuals; with a fine taproot; ± densely pubescent, trichomes several-rayed, rays distinct or fused at base, bifurcate, (prominently tuberculate throughout).
Stems

simple from base, decumbent, (occasionally few-branched), 0.5–1.5(–2) dm.

simple or few to several from base, erect, (sometimes branched), to 4 dm.

Basal leaves

blade elliptic to broadly obovate, triangular, rhombic, or round, 1.5–3(–4) cm, margins often sinuate or shallowly lobed.

blade elliptic to rhombic, 3–8 cm, (base narrowing gradually to petiole), margins entire, repand, coarsely toothed, or pinnatifid.

Cauline leaves

(sessile or shortly petiolate);

blade elliptic to oblanceolate to obovate, 0.5–1.5 cm, (base narrowed to petiole), margins entire.

(proximal often shortly petiolate, distal sessile);

blade linear or narrowly obovate, 1.5–6(–10) cm, margins entire, repand, or shallowly toothed.

Racemes

compact to loose.

usually loose.

Flowers

sepals oblong to broadly elliptic, 4–7.5 mm, (lateral pair saccate or not);

petals spatulate, 7.5–10 mm.

sepals elliptic or ovate, 4–6 mm, (lateral pair usually subsaccate);

petals obovate to obdeltate, 6–10 mm, (apex often emarginate).

Fruiting pedicels

(ascending or divaricate-spreading, straight to loosely sigmoid or curved), 4–10 mm.

(usually divaricate, sometimes horizontal, straight or slightly curved), 8–20 mm.

Fruits

(sessile or substipitate), elliptic, suborbicular, or oblong, strongly compressed (angustiseptate), 5–9 mm, (rounded to sharply keeled on 1 side, edges ± keeled);

valves: (margins covering replum edges or not), usually pubescent throughout or, rarely, glabrous inside;

ovules (4–)8–14(–16) per ovary;

style 2–4.5(–5) mm.

not didymous, ± globose, slightly inflated, 4–6 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4 per ovary;

style 2–3.5 mm; (stigma expanded).

Seeds

slightly flattened.

flattened, (margined).

2n

= 10.

Physaria carinata

Physaria angustifolia

Phenology Flowering Apr(-May).
Habitat Shallow limestone-derived soils, sometimes spreading to disturbed sites
Elevation 90-300 m (300-1000 ft)
Distribution
from FNA
ID; MT; WY
[BONAP county map]
from FNA
OK; TX
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Differences in fruit morphology become blurred and the three subspecies are often indistinguishable where their ranges meet near the intersection of Idaho, Montana, and Wyoming.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits elliptic, not keeled, valve margins (thin, rounded), not covering replum edges.
subsp. paysonii
1. Fruits elliptic, suborbicular, or broadly oblong, keeled on one side, valve margins covering replum edges
→ 2
2. Fruits elliptic, bases narrow-rounded to acute, apices ± acute.
subsp. carinata
2. Fruits suborbicular to elliptic or broadly oblong, bases and apices rounded.
subsp. pulchella
Source FNA vol. 7, p. 629. FNA vol. 7, p. 625.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. carinata subsp. carinata, P. carinata subsp. paysonii, P. carinata subsp. pulchella
Synonyms Lesquerella carinata Vesicaria angustifolia, Lesquerella angustifolia, Lesquerella longifolia
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Nuttall ex Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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