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Calder's bladderpod

Kodachrome bladderpod

Habit Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). Perennials; caudex (buried), branched, (forming hard mats); densely pubescent, trichomes several-rayed, rays furcate or bifurcate, (tuberculate).
Stems

simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm.

several from base, erect, (unbranched), 0.2–0.3 dm.

Basal leaves

blade oblanceolate, 2–3 cm, margins entire.

(few), similar to cauline.

Cauline leaves

(sessile or proximal shortly petiolate);

blade narrowly oblanceolate, 0.5–1.5 cm, margins entire.

(petiole not differentiated from blade);

blade (somewhat succulent), linear to narrowly oblanceolate, 5–12 mm, margins entire.

Racemes

loose.

dense, (few-flowered).

Flowers

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide).

sepals (yellowish), elliptic, 3–4.5 mm;

petals (erect or, more commonly, arching), spatulate to oblanceolate, 5.8–7 mm, (claw not or weakly differentiated from blade).

Fruiting pedicels

(erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout).

(ascending to divaricate-ascending, ± straight), 3.5–6 mm.

Fruits

subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence);

replum as wide as or wider than fruit;

ovules 10–14 per ovary;

style 1–2 mm.

(coppery or reddish brown in age), broadly ovoid, slightly inflated, 3–4 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8 per ovary;

style 1.8–3 mm.

Seeds

plump.

flattened.

2n

= 20.

Physaria calderi

Physaria tumulosa

Phenology Flowering Jun–Aug. Flowering May–Jun.
Habitat Dry rocky summits, limestone flats and slopes, alpine knolls Barren white knolls surrounded by sagebrush, pinyon pine, and Utah juniper
Elevation 600-1500 m (2000-4900 ft) 1600-1800 m (5200-5900 ft)
Distribution
from FNA
AK; NT; YT
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Physaria calderi is known from the Ogilvie and Richardson mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria tumulosa is morphologically similar to 55. P. navajoensis of northeastern Arizona and northwestern New Mexico, and differing very subtly. It has been long treated as an infraspecific taxon of P. hitchcockii; unpublished molecular data do not support that disposition. It is found on knolls of the Winsor Member of the Carmel Formation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 629. FNA vol. 7, p. 664.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella calderi, Lesquerella arctica subsp. calderi Lesquerella hitchcockii subsp. tumulosa, Lesquerella tumulosa, P. rubicundula var. tumulosa
Name authority (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Barneby) O’Kane & Al-Shehbaz: Novon 12: 328. (2002)
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