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Calder's bladderpod

alpine bladderpod, spatula-leaf bladderpod

Habit Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). Perennials; caudex simple, (relatively small); sparsely to moderately pubescent, trichomes 4- or 5-rayed, rays furcate or bifurcate, not fused, (tuberculate).
Stems

simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm.

simple from base, erect to decumbent, (well-exserted beyond basal leaves, loosely spreading), 0.3–1.2 dm.

Basal leaves

blade oblanceolate, 2–3 cm, margins entire.

(erect to prostrate, petiole distinct from blade);

blade (inner) spatulate to oblanceolate, or (outer) oblanceolate or orbicular, 1.5–4 cm, margins entire (rarely folded).

Cauline leaves

(sessile or proximal shortly petiolate);

blade narrowly oblanceolate, 0.5–1.5 cm, margins entire.

blade spatulate, distinctly different from basal.

Racemes

loose.

moderately dense, (6–20-flowered).

Flowers

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide).

sepals (pale yellow), elliptic, 3.5–5 mm;

petals lingulate, 6–9 mm.

Fruiting pedicels

(erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout).

(strongly sigmoid), 10–20 mm (2 times longer than fruits).

Fruits

subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence);

replum as wide as or wider than fruit;

ovules 10–14 per ovary;

style 1–2 mm.

lanceolate or orbicular, slightly inflated, (2.5–)3–6 mm, (apex usually strongly narrowed);

valves pubescent, trichomes sparse and closely appressed to surface;

ovules 4–8 per ovary;

style 2.5–6 mm (usually ± equal in length to mature fruit).

Seeds

plump.

plump.

2n

= 20.

Physaria calderi

Physaria spatulata

Phenology Flowering Jun–Aug. Flowering May-early Jul.
Habitat Dry rocky summits, limestone flats and slopes, alpine knolls Grasslands, subalpine meadows, sagebrush, scattered pines, fellfields, calcareous (sometimes alkaline) substrates
Elevation 600-1500 m (2000-4900 ft) 900-2900 m (3000-9500 ft)
Distribution
from FNA
AK; NT; YT
[BONAP county map]
from FNA
MT; ND; NE; SD; WY; AB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria calderi is known from the Ogilvie and Richardson mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 629. FNA vol. 7, p. 663.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella calderi, Lesquerella arctica subsp. calderi Lesquerella spatulata, Lesquerella alpina var. spatulata, Lesquerella nodosa, P. reediana subsp. spatulata, P. reediana var. spatulata
Name authority (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Rydberg) Grady & O’Kane: Novon 17: 190. (2007)
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