Physaria calderi |
Physaria spatulata |
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Calder's bladderpod |
alpine bladderpod, spatula-leaf bladderpod |
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Habit | Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). | Perennials; caudex simple, (relatively small); sparsely to moderately pubescent, trichomes 4- or 5-rayed, rays furcate or bifurcate, not fused, (tuberculate). |
Stems | simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm. |
simple from base, erect to decumbent, (well-exserted beyond basal leaves, loosely spreading), 0.3–1.2 dm. |
Basal leaves | blade oblanceolate, 2–3 cm, margins entire. |
(erect to prostrate, petiole distinct from blade); blade (inner) spatulate to oblanceolate, or (outer) oblanceolate or orbicular, 1.5–4 cm, margins entire (rarely folded). |
Cauline leaves | (sessile or proximal shortly petiolate); blade narrowly oblanceolate, 0.5–1.5 cm, margins entire. |
blade spatulate, distinctly different from basal. |
Racemes | loose. |
moderately dense, (6–20-flowered). |
Flowers | sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide). |
sepals (pale yellow), elliptic, 3.5–5 mm; petals lingulate, 6–9 mm. |
Fruiting pedicels | (erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout). |
(strongly sigmoid), 10–20 mm (2 times longer than fruits). |
Fruits | subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence); replum as wide as or wider than fruit; ovules 10–14 per ovary; style 1–2 mm. |
lanceolate or orbicular, slightly inflated, (2.5–)3–6 mm, (apex usually strongly narrowed); valves pubescent, trichomes sparse and closely appressed to surface; ovules 4–8 per ovary; style 2.5–6 mm (usually ± equal in length to mature fruit). |
Seeds | plump. |
plump. |
2n | = 20. |
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Physaria calderi |
Physaria spatulata |
|
Phenology | Flowering Jun–Aug. | Flowering May-early Jul. |
Habitat | Dry rocky summits, limestone flats and slopes, alpine knolls | Grasslands, subalpine meadows, sagebrush, scattered pines, fellfields, calcareous (sometimes alkaline) substrates |
Elevation | 600-1500 m (2000-4900 ft) | 900-2900 m (3000-9500 ft) |
Distribution |
AK; NT; YT |
MT; ND; NE; SD; WY; AB; SK
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Discussion | Physaria calderi is known from the Ogilvie and Richardson mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 629. | FNA vol. 7, p. 663. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Lesquerella calderi, Lesquerella arctica subsp. calderi | Lesquerella spatulata, Lesquerella alpina var. spatulata, Lesquerella nodosa, P. reediana subsp. spatulata, P. reediana var. spatulata |
Name authority | (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Rydberg) Grady & O’Kane: Novon 17: 190. (2007) |
Web links |