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Calder's bladderpod

thick-leaf bladderpod

Habit Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). Perennials; caudex usually simple, rarely branched, (loosely mounded, rosette-like growth); densely (silvery or gray) pubescent, trichomes (sessile), 5-rayed, rays bifurcate, slightly fused near base of main rays, (tuberculate throughout, less over umbo).
Stems

simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm.

several from base, decumbent to prostrate, (well-exserted beyond basal leaves), 0.2–0.5 dm.

Basal leaves

blade oblanceolate, 2–3 cm, margins entire.

(petiole differentiated from blade);

blade (slightly cupped, leathery, nearly 1 mm thick), oblanceolate to orbicular, 1.2–2 cm, margins entire, (apex acute).

Cauline leaves

(sessile or proximal shortly petiolate);

blade narrowly oblanceolate, 0.5–1.5 cm, margins entire.

blade spatulate, similar to basal.

Racemes

loose.

dense, (subumbellate).

Flowers

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide).

sepals (pale yellow), elliptic to oblong, 3.5–4.0 mm, (median pair somewhat thickened apically, cucullate);

petals lingulate, 5–6 mm.

Fruiting pedicels

(erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout).

(ascending, curved), 5–7 mm.

Fruits

subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence);

replum as wide as or wider than fruit;

ovules 10–14 per ovary;

style 1–2 mm.

globose or ellipsoid, slightly inflated (with slight apical constriction), 3–6 mm;

valves pubescent, trichomes closely appressed;

ovules 8 per ovary;

style 1–3 mm (shorter than mature fruit).

Seeds

plump.

plump, (oblong).

2n

= 20.

Physaria calderi

Physaria pachyphylla

Phenology Flowering Jun–Aug. Flowering Jun–Jul.
Habitat Dry rocky summits, limestone flats and slopes, alpine knolls Barren areas of mixed white, pink, or reddish limestone and diatomaceous earth
Elevation 600-1500 m (2000-4900 ft) 1300-1600 m (4300-5200 ft)
Distribution
from FNA
AK; NT; YT
[BONAP county map]
from FNA
MT
[BONAP county map]
Discussion

Physaria calderi is known from the Ogilvie and Richardson mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria pachyphylla is known from the Pryor Mountain Desert near the Wyoming state line.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 629. FNA vol. 7, p. 656.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella calderi, Lesquerella arctica subsp. calderi
Name authority (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) O’Kane & Grady: Novon 17: 187, fig. 4. (2007)
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