Physaria calderi |
Physaria ludoviciana |
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Calder's bladderpod |
foothill bladderpod, silver bladderpod |
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Habit | Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). |
Stems | simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm. |
few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
Basal leaves | blade oblanceolate, 2–3 cm, margins entire. |
(erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
Cauline leaves | (sessile or proximal shortly petiolate); blade narrowly oblanceolate, 0.5–1.5 cm, margins entire. |
blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
Racemes | loose. |
compact, (elongated and loose in fruit, densely-flowered). |
Flowers | sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide). |
sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
Fruiting pedicels | (erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout). |
(usually recurved), (5–)10–20(–25) mm. |
Fruits | subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence); replum as wide as or wider than fruit; ovules 10–14 per ovary; style 1–2 mm. |
subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
Seeds | plump. |
slightly flattened. |
2n | = 20. |
= 10, 20, 30. |
Physaria calderi |
Physaria ludoviciana |
|
Phenology | Flowering Jun–Aug. | Flowering Apr–Jul(-Aug). |
Habitat | Dry rocky summits, limestone flats and slopes, alpine knolls | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands |
Elevation | 600-1500 m (2000-4900 ft) | 0-1900 m (0-6200 ft) |
Distribution |
AK; NT; YT |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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Discussion | Physaria calderi is known from the Ogilvie and Richardson mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 629. | FNA vol. 7, p. 649. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Lesquerella calderi, Lesquerella arctica subsp. calderi | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana |
Name authority | (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |