Physaria calderi |
Physaria kingii |
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Calder's bladderpod |
King bladderpod, King's bladder-pod |
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Habit | Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). | Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). | ||||||||||||||||||||||||
Stems | simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm. |
few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm. |
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Basal leaves | blade oblanceolate, 2–3 cm, margins entire. |
blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed. |
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Cauline leaves | (sessile or proximal shortly petiolate); blade narrowly oblanceolate, 0.5–1.5 cm, margins entire. |
(proximal petiolate, distal sessile); blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire. |
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Racemes | loose. |
(usually not secund), dense, (sometimes elongated in fruit). |
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Flowers | sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide). |
sepals lanceolate, 4–6(–7) mm; petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade). |
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Fruiting pedicels | (erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout). |
(erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm. |
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Fruits | subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence); replum as wide as or wider than fruit; ovules 10–14 per ovary; style 1–2 mm. |
(sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute); valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete); ovules 4–16 per ovary; style 1–9 mm. |
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Seeds | plump. |
flattened (sometimes slightly). |
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2n | = 20. |
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Physaria calderi |
Physaria kingii |
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Phenology | Flowering Jun–Aug. | |||||||||||||||||||||||||
Habitat | Dry rocky summits, limestone flats and slopes, alpine knolls | |||||||||||||||||||||||||
Elevation | 600-1500 m (2000-4900 ft) | |||||||||||||||||||||||||
Distribution |
AK; NT; YT |
AZ; CA; ID; NV; OR; UT; nw Mexico
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Discussion | Physaria calderi is known from the Ogilvie and Richardson mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 7 (7 in the flora). The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 629. | FNA vol. 7, p. 645. | ||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||
Synonyms | Lesquerella calderi, Lesquerella arctica subsp. calderi | Vesicaria kingii, Lesquerella kingii | ||||||||||||||||||||||||
Name authority | (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||||||||||||||||||||||
Web links |