Physaria calderi |
Physaria eburniflora |
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Calder's bladderpod |
Devils Gate twinpod |
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Habit | Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). | Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth). |
Stems | simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm. |
simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm. |
Basal leaves | blade oblanceolate, 2–3 cm, margins entire. |
blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed). |
Cauline leaves | (sessile or proximal shortly petiolate); blade narrowly oblanceolate, 0.5–1.5 cm, margins entire. |
(2–4); blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute). |
Racemes | loose. |
condensed. |
Flowers | sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide). |
sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median); petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade). |
Fruiting pedicels | (erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout). |
(divaricate-ascending, nearly straight), 6–10 mm. |
Fruits | subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence); replum as wide as or wider than fruit; ovules 10–14 per ovary; style 1–2 mm. |
strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery); valves (retaining seeds after dehiscence), pubescent; replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse; ovules 4–8 per ovary; style 4–5 mm, (sparsely pubescent or glabrous). |
Seeds | plump. |
plump. |
2n | = 20. |
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Physaria calderi |
Physaria eburniflora |
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Phenology | Flowering Jun–Aug. | Flowering May–Jun. |
Habitat | Dry rocky summits, limestone flats and slopes, alpine knolls | Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus |
Elevation | 600-1500 m (2000-4900 ft) | 1800-3000 m (5900-9800 ft) |
Distribution |
AK; NT; YT |
WY |
Discussion | Physaria calderi is known from the Ogilvie and Richardson mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 629. | FNA vol. 7, p. 635. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Lesquerella calderi, Lesquerella arctica subsp. calderi | |
Name authority | (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | Rollins: Brittonia 33: 333. (1981) |
Web links |