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Calder's bladderpod

Devils Gate twinpod

Habit Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth).
Stems

simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm.

simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm.

Basal leaves

blade oblanceolate, 2–3 cm, margins entire.

blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed).

Cauline leaves

(sessile or proximal shortly petiolate);

blade narrowly oblanceolate, 0.5–1.5 cm, margins entire.

(2–4);

blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute).

Racemes

loose.

condensed.

Flowers

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide).

sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median);

petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade).

Fruiting pedicels

(erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout).

(divaricate-ascending, nearly straight), 6–10 mm.

Fruits

subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence);

replum as wide as or wider than fruit;

ovules 10–14 per ovary;

style 1–2 mm.

strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery);

valves (retaining seeds after dehiscence), pubescent;

replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse;

ovules 4–8 per ovary;

style 4–5 mm, (sparsely pubescent or glabrous).

Seeds

plump.

plump.

2n

= 20.

Physaria calderi

Physaria eburniflora

Phenology Flowering Jun–Aug. Flowering May–Jun.
Habitat Dry rocky summits, limestone flats and slopes, alpine knolls Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus
Elevation 600-1500 m (2000-4900 ft) 1800-3000 m (5900-9800 ft)
Distribution
from FNA
AK; NT; YT
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Physaria calderi is known from the Ogilvie and Richardson mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 629. FNA vol. 7, p. 635.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella calderi, Lesquerella arctica subsp. calderi
Name authority (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) Rollins: Brittonia 33: 333. (1981)
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