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Calder's bladderpod

Great Plains bladderpod

Habit Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). Perennials or, rarely, annuals; caudex simple or branched; ± densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (usually spreading), distinct or slightly fused at base, furcate or bifurcate, (tuberculate).
Stems

simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm.

simple or few from base, prostrate or straggling to erect, (sometimes purplish, usually unbranched), (0.5–)1–2(–3) dm.

Basal leaves

blade oblanceolate, 2–3 cm, margins entire.

blade oblanceolate, 1.5–5(–7) cm, margins entire or shallowly dentate, (flat).

Cauline leaves

(sessile or proximal shortly petiolate);

blade narrowly oblanceolate, 0.5–1.5 cm, margins entire.

blade elliptic to linear, (0.5–)1–2.5(–3) cm, margins usually entire.

Racemes

loose.

(secund), dense, (elongated in fruit).

Flowers

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide).

sepals elliptic or oblong, 4–6(–7) mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals (often red or lavender when dried), obovate, 6–8.5(–9.5) mm, (narrowing to broad claw).

Fruiting pedicels

(erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout).

(usually sharply recurved, sometimes divaricate-spreading or nearly horizontal), 5–15(–20) mm, (stout).

Fruits

subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence);

replum as wide as or wider than fruit;

ovules 10–14 per ovary;

style 1–2 mm.

subglobose, obovoid, or broadly ellipsoid, slightly inflated, (3.5–)4–5.5(–6.5) mm;

valves densely pubescent outside, trichomes spreading or closely appressed, rarely sparsely pubescent inside;

ovules (4–)8(–10) per ovary;

style (slender), 3–5.5(–6.5) mm.

Seeds

plump.

slightly flattened.

2n

= 20.

Physaria calderi

Physaria arenosa

Phenology Flowering Jun–Aug.
Habitat Dry rocky summits, limestone flats and slopes, alpine knolls
Elevation 600-1500 m (2000-4900 ft)
Distribution
from FNA
AK; NT; YT
[BONAP county map]
from FNA
CO; MT; ND; NE; SD; WY; AB; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria calderi is known from the Ogilvie and Richardson mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perennials or, rarely, annuals, short-lived; fruit valves: trichomes spreading.
subsp. arenosa
1. Perennials, long-lived; fruit valves: trichomes closely appressed.
subsp. argillosa
Source FNA vol. 7, p. 629. FNA vol. 7, p. 626.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. arenosa subsp. arenosa, P. arenosa subsp. argillosa
Synonyms Lesquerella calderi, Lesquerella arctica subsp. calderi Vesicaria arenosa, Lesquerella arenosa, Lesquerella argentea var. arenosa, Lesquerella ludoviciana var. arenosa
Name authority (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Richardson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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