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Rocky Mountain bladderpod

Navajo bladderpod

Habit Perennials; (compact); caudex branched; densely (silvery) pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays distinct, furcate or bifurcate, (umbonate, tuberculate and the center less so). Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo).
Stems

several from base, erect or outer ones decumbent, (unbranched, stout, usually sparsely leaved), 1–3 dm.

several from base (crowded), erect, not exceeding leaves.

Basal leaves

blade linear, 2–7(–10) cm, margins entire, repand, or shallowly dentate.

usually absent.

Cauline leaves

(sessile);

blade (erect), spatulate to linear, (1–)2–3(–4.5) cm, margins entire, sometimes involute, (apex acute or subacute).

(petiole not differentiated from blade);

blade linear-oblanceolate, 3–8(–13) mm, margins entire.

Racemes

dense, (exceeding basal leaves).

(secund), dense, corymbose, (few-flowered, not or barely exceeding leaves).

Flowers

sepals ovate or oblong, (4.5–)5–6(–7) mm, (lateral pair subsaccate, cucullate, median pair thickened, cucullate apically);

petals spatulate, 7–9(–11) mm (widened at base, slightly retuse).

sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate);

petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle).

Fruiting pedicels

(spreading, sharply sigmoid), 8–15 mm.

(ascending to divaricate-ascending, straight), 3.5–6 mm.

Fruits

(sessile or substipitate), ovate to oblong, not compressed at distal margins or apex, 5–9 mm;

valves sparsely pubescent, trichomes appressed;

ovules 4–8 per ovary;

style 3–5 mm.

(becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute);

valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not);

ovules 4–8 per ovary;

style 1.8–3 mm.

Seeds

flattened.

plump or slightly flattened, (strongly mucilaginous).

2n

= 16, ca. 20.

Physaria calcicola

Physaria navajoensis

Phenology Flowering May–Jun. Flowering May-early Jun.
Habitat Shale bluffs, limestone hillsides, gypseous knolls and ravines, calcareous substrates, grasslands and pinyon-juniper communities Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone
Elevation 1400-2100 m (4600-6900 ft) 2200-2400 m (7200-7900 ft)
Distribution
from FNA
CO; NM
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 629. FNA vol. 7, p. 651.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella calcicola Lesquerella navajoensis
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (O’Kane) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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