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Rocky Mountain bladderpod

Hitchcock's bladderpod

Habit Perennials; (compact); caudex branched; densely (silvery) pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays distinct, furcate or bifurcate, (umbonate, tuberculate and the center less so). Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate).
Stems

several from base, erect or outer ones decumbent, (unbranched, stout, usually sparsely leaved), 1–3 dm.

few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm.

Basal leaves

blade linear, 2–7(–10) cm, margins entire, repand, or shallowly dentate.

(petiole and blade differentiated or not);

blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire.

Cauline leaves

(sessile);

blade (erect), spatulate to linear, (1–)2–3(–4.5) cm, margins entire, sometimes involute, (apex acute or subacute).

similar to basal, smaller.

Racemes

dense, (exceeding basal leaves).

dense.

Flowers

sepals ovate or oblong, (4.5–)5–6(–7) mm, (lateral pair subsaccate, cucullate, median pair thickened, cucullate apically);

petals spatulate, 7–9(–11) mm (widened at base, slightly retuse).

sepals narrowly lanceolate to lanceolate, 2.8–6 mm;

petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade).

Fruiting pedicels

(spreading, sharply sigmoid), 8–15 mm.

(ascending, straight or slightly curved), 2–6 mm.

Fruits

(sessile or substipitate), ovate to oblong, not compressed at distal margins or apex, 5–9 mm;

valves sparsely pubescent, trichomes appressed;

ovules 4–8 per ovary;

style 3–5 mm.

(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute);

valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8 per ovary;

style 1.7–6 mm.

Seeds

flattened.

flattened.

2n

= 16, ca. 20.

Physaria calcicola

Physaria hitchcockii

Phenology Flowering May–Jun.
Habitat Shale bluffs, limestone hillsides, gypseous knolls and ravines, calcareous substrates, grasslands and pinyon-juniper communities
Elevation 1400-2100 m (4600-6900 ft)
Distribution
from FNA
CO; NM
[WildflowerSearch map]
[BONAP county map]
from FNA
NV; UT
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices elongated and elastic; basal leaves: petiole not differentiated from blade, blades linear-oblanceolate; Aquarius, Markagunt, and Paunsaugunt plateaus, Utah (limited to the pink member of the limestone Wasatch (Claron) Formation).
subsp. rubicundula
1. Caudices elongated or not, elastic or not; basal leaves: petiole differentiated (sometimes weakly) from blade, blades oblanceolate to obovate; Nevada, Utah
→ 2
2. Plants forming tufts; caudices not elongated, not elastic; fruits 2.6-3.8 mm wide; Table Cliff Plateau, Utah (limited to the white member of the limestone Wasatch (Claron) Formation) and limestones of the Sheep Range and Spring Mountains, Nevada.
subsp. hitchcockii
2. Plants forming soft mats; caudices elongated, elastic (creeping); fruits 1.7-3 mm wide; Grant, Quinn Canyon, and Schell Creek ranges, Nevada.
subsp. confluens
Source FNA vol. 7, p. 629. FNA vol. 7, p. 643.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. hitchcockii subsp. confluens, P. hitchcockii subsp. hitchcockii, P. hitchcockii subsp. rubicundula
Synonyms Lesquerella calcicola Lesquerella hitchcockii
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002)
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