Physaria calcicola |
Physaria grahamii |
|
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Rocky Mountain bladderpod |
Graham's twinpod |
|
Habit | Perennials; (compact); caudex branched; densely (silvery) pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays distinct, furcate or bifurcate, (umbonate, tuberculate and the center less so). | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. |
Stems | several from base, erect or outer ones decumbent, (unbranched, stout, usually sparsely leaved), 1–3 dm. |
several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
Basal leaves | blade linear, 2–7(–10) cm, margins entire, repand, or shallowly dentate. |
(outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
Cauline leaves | (sessile); blade (erect), spatulate to linear, (1–)2–3(–4.5) cm, margins entire, sometimes involute, (apex acute or subacute). |
similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
Racemes | dense, (exceeding basal leaves). |
loose, (elongated). |
Flowers | sepals ovate or oblong, (4.5–)5–6(–7) mm, (lateral pair subsaccate, cucullate, median pair thickened, cucullate apically); petals spatulate, 7–9(–11) mm (widened at base, slightly retuse). |
sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
Fruiting pedicels | (spreading, sharply sigmoid), 8–15 mm. |
(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
Fruits | (sessile or substipitate), ovate to oblong, not compressed at distal margins or apex, 5–9 mm; valves sparsely pubescent, trichomes appressed; ovules 4–8 per ovary; style 3–5 mm. |
didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
Seeds | flattened. |
plump, (suborbicular). |
2n | = 16, ca. 20. |
|
Physaria calcicola |
Physaria grahamii |
|
Phenology | Flowering May–Jun. | Flowering May–Jun. |
Habitat | Shale bluffs, limestone hillsides, gypseous knolls and ravines, calcareous substrates, grasslands and pinyon-juniper communities | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay |
Elevation | 1400-2100 m (4600-6900 ft) | 2100-2900 m (6900-9500 ft) |
Distribution |
CO; NM
|
UT |
Discussion | Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 629. | FNA vol. 7, p. 642. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella calcicola | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda |
Name authority | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) |
Web links |