Physaria calcicola |
Physaria douglasii |
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Rocky Mountain bladderpod |
Columbia bladder-pod, Douglas' bladder-pod |
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Habit | Perennials; (compact); caudex branched; densely (silvery) pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays distinct, furcate or bifurcate, (umbonate, tuberculate and the center less so). | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | ||||
Stems | several from base, erect or outer ones decumbent, (unbranched, stout, usually sparsely leaved), 1–3 dm. |
simple from base, erect, (usually unbranched), to 4.5 dm. |
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Basal leaves | blade linear, 2–7(–10) cm, margins entire, repand, or shallowly dentate. |
blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
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Cauline leaves | (sessile); blade (erect), spatulate to linear, (1–)2–3(–4.5) cm, margins entire, sometimes involute, (apex acute or subacute). |
similar to basal, blade narrowly linear or, sometimes, orbicular. |
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Racemes | dense, (exceeding basal leaves). |
loose (lax). |
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Flowers | sepals ovate or oblong, (4.5–)5–6(–7) mm, (lateral pair subsaccate, cucullate, median pair thickened, cucullate apically); petals spatulate, 7–9(–11) mm (widened at base, slightly retuse). |
sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
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Fruiting pedicels | (spreading, sharply sigmoid), 8–15 mm. |
(recurved, straight, curved, or sigmoid), 6–20 mm. |
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Fruits | (sessile or substipitate), ovate to oblong, not compressed at distal margins or apex, 5–9 mm; valves sparsely pubescent, trichomes appressed; ovules 4–8 per ovary; style 3–5 mm. |
obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 16, ca. 20. |
= 10, 30. |
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Physaria calcicola |
Physaria douglasii |
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Phenology | Flowering May–Jun. | |||||
Habitat | Shale bluffs, limestone hillsides, gypseous knolls and ravines, calcareous substrates, grasslands and pinyon-juniper communities | |||||
Elevation | 1400-2100 m (4600-6900 ft) | |||||
Distribution |
CO; NM
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ID; MT; OR; WA; BC
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Discussion | Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 629. | FNA vol. 7, p. 634. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella calcicola | Lesquerella douglasii | ||||
Name authority | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
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