Physaria calcicola |
Physaria didymocarpa |
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Rocky Mountain bladderpod |
common twinpod, double twinpod, twin bladderpod |
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Habit | Perennials; (compact); caudex branched; densely (silvery) pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays distinct, furcate or bifurcate, (umbonate, tuberculate and the center less so). | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | ||||||||
Stems | several from base, erect or outer ones decumbent, (unbranched, stout, usually sparsely leaved), 1–3 dm. |
several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
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Basal leaves | blade linear, 2–7(–10) cm, margins entire, repand, or shallowly dentate. |
(forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
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Cauline leaves | (sessile); blade (erect), spatulate to linear, (1–)2–3(–4.5) cm, margins entire, sometimes involute, (apex acute or subacute). |
blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
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Racemes | dense, (exceeding basal leaves). |
congested, (elongated in fruit, greatly exceeding leaves). |
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Flowers | sepals ovate or oblong, (4.5–)5–6(–7) mm, (lateral pair subsaccate, cucullate, median pair thickened, cucullate apically); petals spatulate, 7–9(–11) mm (widened at base, slightly retuse). |
sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
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Fruiting pedicels | (spreading, sharply sigmoid), 8–15 mm. |
(spreading, straight or slightly curved), 8–12 mm. |
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Fruits | (sessile or substipitate), ovate to oblong, not compressed at distal margins or apex, 5–9 mm; valves sparsely pubescent, trichomes appressed; ovules 4–8 per ovary; style 3–5 mm. |
(erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 16, ca. 20. |
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Physaria calcicola |
Physaria didymocarpa |
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Phenology | Flowering May–Jun. | |||||||||
Habitat | Shale bluffs, limestone hillsides, gypseous knolls and ravines, calcareous substrates, grasslands and pinyon-juniper communities | |||||||||
Elevation | 1400-2100 m (4600-6900 ft) | |||||||||
Distribution |
CO; NM
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ID; MT; WA; WY; AB; BC
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Discussion | Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 629. | FNA vol. 7, p. 633. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lesquerella calcicola | Vesicaria didymocarpa, Coulterina didymocarpa | ||||||||
Name authority | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | ||||||||
Web links |