Physaria calcicola |
Physaria acutifolia |
|
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Rocky Mountain bladderpod |
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
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Habit | Perennials; (compact); caudex branched; densely (silvery) pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays distinct, furcate or bifurcate, (umbonate, tuberculate and the center less so). | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). |
Stems | several from base, erect or outer ones decumbent, (unbranched, stout, usually sparsely leaved), 1–3 dm. |
several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
Basal leaves | blade linear, 2–7(–10) cm, margins entire, repand, or shallowly dentate. |
(petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
Cauline leaves | (sessile); blade (erect), spatulate to linear, (1–)2–3(–4.5) cm, margins entire, sometimes involute, (apex acute or subacute). |
blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
Racemes | dense, (exceeding basal leaves). |
loose, (elongated in fruit). |
Flowers | sepals ovate or oblong, (4.5–)5–6(–7) mm, (lateral pair subsaccate, cucullate, median pair thickened, cucullate apically); petals spatulate, 7–9(–11) mm (widened at base, slightly retuse). |
sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
Fruiting pedicels | (spreading, sharply sigmoid), 8–15 mm. |
(divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
Fruits | (sessile or substipitate), ovate to oblong, not compressed at distal margins or apex, 5–9 mm; valves sparsely pubescent, trichomes appressed; ovules 4–8 per ovary; style 3–5 mm. |
(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
Seeds | flattened. |
(dark brown), flattened, (2–3 mm). |
2n | = 16, ca. 20. |
= 10, 16, 24. |
Physaria calcicola |
Physaria acutifolia |
|
Phenology | Flowering May–Jun. | Flowering May–Jun(-Jul). |
Habitat | Shale bluffs, limestone hillsides, gypseous knolls and ravines, calcareous substrates, grasslands and pinyon-juniper communities | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities |
Elevation | 1400-2100 m (4600-6900 ft) | 1500-3500 m (4900-11500 ft) |
Distribution |
CO; NM
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AZ; CO; ID; MT; NM; NV; SD; UT; WY
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Discussion | Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 629. | FNA vol. 7, p. 624. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella calcicola | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa |
Name authority | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) |
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