Physaria brassicoides |
Physaria scrotiformis |
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double bladderpod, double twinpod |
silver twinpod, west silver bladderpod |
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Habit | Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout). | Perennials; (diminutive); caudex simple or branched, (buried, with thatch of persistent leaf bases distally); (appearing silvery gray-green to silvery purple), densely pubescent, trichomes usually 5 or 6 (rarely 7)-rayed, rays bifurcate or incompletely so, (relatively short, stout, umbonate, moderately tuberculate to nearly smooth, lower layer smoother). |
Stems | several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm. |
1–5 from base, prostrate to slightly decumbent, (arising laterally, also erect or ascending from tuft of basal leaves, unbranched, purple-green), 0.08–0.3 dm. |
Basal leaves | (petiole somewhat winged); blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy). |
(petiole slightly winged); blade oblanceolate, elliptic, or rhombic, (mostly flat, sometimes somewhat folded), 0.6–2.7 cm, (base tapering to petiole), margins entire, (apex rounded to rounded-acute). |
Cauline leaves | blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute). |
(3–7, shortly petiolate or sessile); blade elliptic to oblanceolate, 0.3–0.5 cm, margins entire. |
Racemes | moderately dense (or elongated). |
crowded, (ca. 3–7 fruits). |
Flowers | sepals linear-oblong, 6–8 mm; petals spatulate, 9–12 mm. |
sepals (greenish yellow), linear-triangular, 3.7–5 mm, (lateral pair subsaccate); petals oblanceolate to narrowly obovate, 4.5–9 mm. |
Fruiting pedicels | (divergent, straight to somewhat curved or sigmoid), 5–12 mm. |
(ascending, straight), 1.8–3.4 mm. |
Fruits | (erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad); valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading; replum linear-oblong, constricted, as wide as or wider than fruit; ovules 4 per ovary; style 4–5(–9) mm. |
(shortly stipitate, purple or greenish purple in age), slightly didymous, ovoid to obpyriform, 3–5 mm (wider than long, base rounded-obtuse, apex rounded, flattened, or slightly emarginate); valves (inflated, slightly wider than replum), pubescent, trichomes scattered; replum obovate to orbicular-obdeltate, apex rounded, obtuse, or truncate; septum complete or medially small-perforate; ovules 4–6(–8) per ovary; style 2–3.6 mm. |
Seeds | plump, (broad). |
relatively plump, (ovate to suborbicular, usually rounded on one side, ± flat or concave on the other, not mucilaginous when wetted). |
2n | = 8, 16. |
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Physaria brassicoides |
Physaria scrotiformis |
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Phenology | Flowering May–Jun. | Flowering Jun-early Jul. |
Habitat | Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks | Tundra areas with islands of Engelmann spruce on Leadville limestone, amidst limestone cobbles and gravel |
Elevation | 900-1400 m (3000-4600 ft) | 3500-3700 m (11500-12100 ft) |
Distribution |
CO; MT; ND; NE; SD; WY
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CO |
Discussion | Physaria scrotiformis is known only from La Plata County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 662. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 237. (1902) | O’Kane: Novon 17: 376, fig. 1. (2007) |
Web links |