Physaria brassicoides |
Physaria grahamii |
|
---|---|---|
double bladderpod, double twinpod |
Graham's twinpod |
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Habit | Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout). | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. |
Stems | several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm. |
several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
Basal leaves | (petiole somewhat winged); blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy). |
(outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
Cauline leaves | blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute). |
similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
Racemes | moderately dense (or elongated). |
loose, (elongated). |
Flowers | sepals linear-oblong, 6–8 mm; petals spatulate, 9–12 mm. |
sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
Fruiting pedicels | (divergent, straight to somewhat curved or sigmoid), 5–12 mm. |
(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
Fruits | (erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad); valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading; replum linear-oblong, constricted, as wide as or wider than fruit; ovules 4 per ovary; style 4–5(–9) mm. |
didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
Seeds | plump, (broad). |
plump, (suborbicular). |
2n | = 8, 16. |
|
Physaria brassicoides |
Physaria grahamii |
|
Phenology | Flowering May–Jun. | Flowering May–Jun. |
Habitat | Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay |
Elevation | 900-1400 m (3000-4600 ft) | 2100-2900 m (6900-9500 ft) |
Distribution |
CO; MT; ND; NE; SD; WY
|
UT |
Discussion | Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 642. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda | |
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 237. (1902) | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) |
Web links |