The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

double bladderpod, double twinpod

spreading bladderpod

Habit Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout). Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate).
Stems

several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm.

simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm.

Basal leaves

(petiole somewhat winged);

blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy).

blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent).

Cauline leaves

blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute).

(proximal petiolate, distal sessile);

blade oblanceolate to oblong, 1–7 cm, margins dentate to repand.

Racemes

moderately dense (or elongated).

loose, (elongated).

Flowers

sepals linear-oblong, 6–8 mm;

petals spatulate, 9–12 mm.

sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate);

petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw).

Fruiting pedicels

(divergent, straight to somewhat curved or sigmoid), 5–12 mm.

(usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout).

Fruits

(erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad);

valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading;

replum linear-oblong, constricted, as wide as or wider than fruit;

ovules 4 per ovary;

style 4–5(–9) mm.

(stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm;

valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside;

replum as wide as or wider than fruit;

ovules 8–20(–28) per ovary;

style 2–4.5 mm.

Seeds

plump, (broad).

slightly flattened.

2n

= 8, 16.

Physaria brassicoides

Physaria gracilis

Phenology Flowering May–Jun.
Habitat Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks
Elevation 900-1400 m (3000-4600 ft)
Distribution
from FNA
CO; MT; ND; NE; SD; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems to 7 dm; cauline leaves: blade margins usually deeply dentate, rarely repand; fruits ± sessile, globose or subglobose, 3-6 mm, bases rounded.
subsp. gracilis
1. Stems usually less than 3 dm; cauline leaves: blade margins frequently repand, occasionally dentate; fruits stipitate (gynophore slender), obpyriform to narrowly obovoid, (5-)6-9 mm, bases truncate.
subsp. nuttallii
Source FNA vol. 7, p. 628. FNA vol. 7, p. 641.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. gracilis subsp. gracilis, P. gracilis subsp. nuttallii
Synonyms Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis
Name authority Rydberg: Bull. Torrey Bot. Club 29: 237. (1902) (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
Web links