Physaria brassicoides |
Physaria gracilis |
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double bladderpod, double twinpod |
spreading bladderpod |
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Habit | Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout). | Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). | ||||
Stems | several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm. |
simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. |
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Basal leaves | (petiole somewhat winged); blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy). |
blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). |
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Cauline leaves | blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute). |
(proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. |
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Racemes | moderately dense (or elongated). |
loose, (elongated). |
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Flowers | sepals linear-oblong, 6–8 mm; petals spatulate, 9–12 mm. |
sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). |
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Fruiting pedicels | (divergent, straight to somewhat curved or sigmoid), 5–12 mm. |
(usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). |
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Fruits | (erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad); valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading; replum linear-oblong, constricted, as wide as or wider than fruit; ovules 4 per ovary; style 4–5(–9) mm. |
(stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. |
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Seeds | plump, (broad). |
slightly flattened. |
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2n | = 8, 16. |
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Physaria brassicoides |
Physaria gracilis |
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Phenology | Flowering May–Jun. | |||||
Habitat | Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks | |||||
Elevation | 900-1400 m (3000-4600 ft) | |||||
Distribution |
CO; MT; ND; NE; SD; WY
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AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 641. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis | |||||
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 237. (1902) | (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |