Physaria brassicoides |
Physaria chambersii |
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double bladderpod, double twinpod |
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
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Habit | Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout). | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). |
Stems | several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm. |
several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
Basal leaves | (petiole somewhat winged); blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy). |
(petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
Cauline leaves | blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute). |
blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
Racemes | moderately dense (or elongated). |
congested. |
Flowers | sepals linear-oblong, 6–8 mm; petals spatulate, 9–12 mm. |
sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
Fruiting pedicels | (divergent, straight to somewhat curved or sigmoid), 5–12 mm. |
(divaricate, slightly sigmoid), 8–15 mm. |
Fruits | (erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad); valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading; replum linear-oblong, constricted, as wide as or wider than fruit; ovules 4 per ovary; style 4–5(–9) mm. |
(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
Seeds | plump, (broad). |
flattened. |
2n | = 8, 16. |
= 8, 10, 16, 24. |
Physaria brassicoides |
Physaria chambersii |
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Phenology | Flowering May–Jun. | Flowering Apr–Jul. |
Habitat | Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas |
Elevation | 900-1400 m (3000-4600 ft) | 1500-3200 m (4900-10500 ft) |
Distribution |
CO; MT; ND; NE; SD; WY
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AZ; CA; ID; NV; OR; UT
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 631. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 237. (1902) | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) |
Web links |