Physaria brassicoides |
Physaria carinata |
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double bladderpod, double twinpod |
Idaho bladderpod |
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Habit | Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout). | Perennials; caudex simple, (often enlarged by persistent leaf bases); densely pubescent, trichomes (sessile or short-stalked), rays furcate or bifurcate, (nearly smooth to finely tuberculate). | ||||||||
Stems | several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm. |
simple from base, decumbent, (occasionally few-branched), 0.5–1.5(–2) dm. |
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Basal leaves | (petiole somewhat winged); blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy). |
blade elliptic to broadly obovate, triangular, rhombic, or round, 1.5–3(–4) cm, margins often sinuate or shallowly lobed. |
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Cauline leaves | blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute). |
(sessile or shortly petiolate); blade elliptic to oblanceolate to obovate, 0.5–1.5 cm, (base narrowed to petiole), margins entire. |
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Racemes | moderately dense (or elongated). |
compact to loose. |
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Flowers | sepals linear-oblong, 6–8 mm; petals spatulate, 9–12 mm. |
sepals oblong to broadly elliptic, 4–7.5 mm, (lateral pair saccate or not); petals spatulate, 7.5–10 mm. |
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Fruiting pedicels | (divergent, straight to somewhat curved or sigmoid), 5–12 mm. |
(ascending or divaricate-spreading, straight to loosely sigmoid or curved), 4–10 mm. |
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Fruits | (erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad); valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading; replum linear-oblong, constricted, as wide as or wider than fruit; ovules 4 per ovary; style 4–5(–9) mm. |
(sessile or substipitate), elliptic, suborbicular, or oblong, strongly compressed (angustiseptate), 5–9 mm, (rounded to sharply keeled on 1 side, edges ± keeled); valves: (margins covering replum edges or not), usually pubescent throughout or, rarely, glabrous inside; ovules (4–)8–14(–16) per ovary; style 2–4.5(–5) mm. |
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Seeds | plump, (broad). |
slightly flattened. |
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2n | = 8, 16. |
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Physaria brassicoides |
Physaria carinata |
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Phenology | Flowering May–Jun. | |||||||||
Habitat | Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks | |||||||||
Elevation | 900-1400 m (3000-4600 ft) | |||||||||
Distribution |
CO; MT; ND; NE; SD; WY
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ID; MT; WY |
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Discussion | Subspecies 3 (3 in the flora). Differences in fruit morphology become blurred and the three subspecies are often indistinguishable where their ranges meet near the intersection of Idaho, Montana, and Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 629. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lesquerella carinata | |||||||||
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 237. (1902) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||||||
Web links |