Physaria brassicoides |
Physaria arctica |
|
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double bladderpod, double twinpod |
arctic bladderpod |
|
Habit | Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout). | Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth). |
Stems | several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm. |
simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm. |
Basal leaves | (petiole somewhat winged); blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy). |
(usually ± rosulate); blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire. |
Cauline leaves | blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute). |
(sessile or shortly petiolate); blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire. |
Racemes | moderately dense (or elongated). |
loose. |
Flowers | sepals linear-oblong, 6–8 mm; petals spatulate, 9–12 mm. |
sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw). |
Fruiting pedicels | (divergent, straight to somewhat curved or sigmoid), 5–12 mm. |
(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout). |
Fruits | (erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad); valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading; replum linear-oblong, constricted, as wide as or wider than fruit; ovules 4 per ovary; style 4–5(–9) mm. |
subglobose to ellipsoid, uncompressed, 4–6(–9) mm; valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile; replum as wide as or wider than fruit; ovules (8–)10–14(–16) per ovary; style 1–2.5(–4) mm. |
Seeds | plump, (broad). |
plump. |
2n | = 8, 16. |
= 60. |
Physaria brassicoides |
Physaria arctica |
|
Phenology | Flowering May–Jun. | Flowering May–Aug. |
Habitat | Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks | Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance |
Elevation | 900-1400 m (3000-4600 ft) | 0-1800 m (0-5900 ft) |
Distribution |
CO; MT; ND; NE; SD; WY
|
AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia) |
Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 626. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa | |
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 237. (1902) | (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
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