Physaria brassicoides |
Physaria acutifolia |
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double bladderpod, double twinpod |
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
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Habit | Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout). | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). |
Stems | several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm. |
several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
Basal leaves | (petiole somewhat winged); blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy). |
(petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
Cauline leaves | blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute). |
blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
Racemes | moderately dense (or elongated). |
loose, (elongated in fruit). |
Flowers | sepals linear-oblong, 6–8 mm; petals spatulate, 9–12 mm. |
sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
Fruiting pedicels | (divergent, straight to somewhat curved or sigmoid), 5–12 mm. |
(divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
Fruits | (erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad); valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading; replum linear-oblong, constricted, as wide as or wider than fruit; ovules 4 per ovary; style 4–5(–9) mm. |
(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
Seeds | plump, (broad). |
(dark brown), flattened, (2–3 mm). |
2n | = 8, 16. |
= 10, 16, 24. |
Physaria brassicoides |
Physaria acutifolia |
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Phenology | Flowering May–Jun. | Flowering May–Jun(-Jul). |
Habitat | Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities |
Elevation | 900-1400 m (3000-4600 ft) | 1500-3500 m (4900-11500 ft) |
Distribution |
CO; MT; ND; NE; SD; WY
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AZ; CO; ID; MT; NM; NV; SD; UT; WY
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Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 624. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | |
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 237. (1902) | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) |
Web links |