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Bell's or Front Range twinpod, Bell's twinpod, Front Range twinpod

Nelson's bladderpod

Habit Perennials; caudex simple, (relatively large); densely (silvery) pubescent, trichomes (sessile, appressed), rays furcate, fused at base. Perennials; (diminutive); caudex branched, (densely cespitose, mound-forming); densely pubescent, trichomes (flaring, giving a shaggy appearance), usually 4–5-rayed, rays furcate or bifurcate, (strongly tuberculate throughout).
Stems

simple from base, decumbent to nearly prostrate, 0.5–1.3 dm.

few to several from base, erect, (from basal tuft), 0.1–0.2(–0.3) dm, (not or just barely exceeding leaves).

Basal leaves

(strongly rosulate; shortly petiolate);

blade broadly obovate, 1.5–7.5 (width 7.5–26 mm, base gradually tapering to petiole), margins shallowly dentate, (apex obtuse).

blade spatulate to oblanceolate, 0.5–1.5 cm, (base gradually narrowed to petiole), margins entire.

Cauline leaves

blade oblanceolate to broadly obovate, 1–2.5 cm, margins entire.

(absent or few);

similar to basal, blade linear.

Racemes

dense.

dense, (few-flowered).

Flowers

sepals (pale yellow or yellow-green), narrowly lanceolate to narrowly deltate, 4–8 mm;

petals yellow, broadly spatulate to obovate, 9–13 mm, (not clawed).

sepals (pale yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate);

petals (bright yellow), lingulate, 6–8 mm.

Fruiting pedicels

(divaricate-ascending to widely spreading, slightly sigmoid to curved), 7–12 mm.

(loosely sigmoid), 3–5 mm.

Fruits

didymous, slightly flattened (contrary to replum) to uncompressed, 4–9 × 2–8 mm, (strongly coriaceous, apical and basal sinuses narrow, deep);

valves (retaining seeds after dehiscence), pubescent, trichomes appressed;

replum narrowly oblanceolate to narrowly linear-oblong, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style more than 3 mm.

lanceolate, compressed apically, 2.5–3(–4) mm;

valves pubescent, trichomes spreading, (appearing fuzzy), rarely with trichomes inside;

ovules 4–8 per ovary;

style 2.5–4 mm.

Seeds

compressed.

plump, (oblong).

2n

= 8.

Physaria bellii

Physaria nelsonii

Phenology Flowering Mar–Jun(-Jul). Flowering May-mid Jun.
Habitat Dark shale, road cuts, ridge crests, washes Limestone, windswept knolls and cliffs, nearly barren areas with other cushion-forming plants
Elevation 1500-1800 m (4900-5900 ft) 1600-2300 m (5200-7500 ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
UT; WY
[BONAP county map]
Discussion

Of conservation concern.

Physaria bellii is often found in shale and limestone soils of the Fountain/Ingleside, Lykins, Niobrara, and Pierre formations. It is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Physaria nelsonii is morphologically similar to 73. P. pycnantha, which traditionally was included in a broader P. nelsonii. These allopatric species are distinguished by styles equal to or exceeding the length of fruit and fruiting stems overtopped by basal leaves (P. nelsonii) versus styles shorter than fruits and fruiting stems usually exserted beyond basal leaves (P. pycnantha).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 628. FNA vol. 7, p. 652.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella condensata, Lesquerella alpina subsp. condensata, Lesquerella alpina var. condensata
Name authority G. A. Mulligan: Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. (1966) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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