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Bell's or Front Range twinpod, Bell's twinpod, Front Range twinpod

Mcvaugh's bladderpod

Habit Perennials; caudex simple, (relatively large); densely (silvery) pubescent, trichomes (sessile, appressed), rays furcate, fused at base. Perennials; caudex simple or branched, (sometimes enlarged); densely pubescent, trichomes (sessile), several-rayed, rays fused (webbed) most of their length, (umbonate, peltate, tuberculate throughout).
Stems

simple from base, decumbent to nearly prostrate, 0.5–1.3 dm.

few to several from base, erect or outer ones decumbent, 0.5–4 dm.

Basal leaves

(strongly rosulate; shortly petiolate);

blade broadly obovate, 1.5–7.5 (width 7.5–26 mm, base gradually tapering to petiole), margins shallowly dentate, (apex obtuse).

(long-petiolate);

blade elliptic to obovate or rhombic, 2–6(–9) cm, margins entire.

Cauline leaves

blade oblanceolate to broadly obovate, 1–2.5 cm, margins entire.

(sessile or shortly petiolate);

blade oblanceolate to spatulate, 1–3 cm, (proximal broader), margins entire.

Racemes

dense.

dense, (relatively short).

Flowers

sepals (pale yellow or yellow-green), narrowly lanceolate to narrowly deltate, 4–8 mm;

petals yellow, broadly spatulate to obovate, 9–13 mm, (not clawed).

sepals elliptic or narrowly oblong, 4–5.4 mm, (tapered to apex);

petals (white, base and claw yellow, conspicuously purple-veined), usually broadly obovate or rhombic, 6–10 mm, (± equal to blade, tapering to slender claw).

Fruiting pedicels

(divaricate-ascending to widely spreading, slightly sigmoid to curved), 7–12 mm.

(erect to spreading, ascending, or (proximal) horizontal, straight to slightly curved, sometimes loosely sigmoid), 6–12(–20) mm.

Fruits

didymous, slightly flattened (contrary to replum) to uncompressed, 4–9 × 2–8 mm, (strongly coriaceous, apical and basal sinuses narrow, deep);

valves (retaining seeds after dehiscence), pubescent, trichomes appressed;

replum narrowly oblanceolate to narrowly linear-oblong, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style more than 3 mm.

(sessile or substipitate, often reddish magenta), usually ovoid to subglobose, inflated, 4–6(–7) mm;

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

septum perforate;

ovules 8–12 per ovary;

style 1.5–4 mm.

Seeds

compressed.

somewhat flattened.

2n

= 8.

= 12.

Physaria bellii

Physaria mcvaughiana

Phenology Flowering Mar–Jun(-Jul). Flowering mid Mar–Apr(-Aug).
Habitat Dark shale, road cuts, ridge crests, washes Stream bed gravels, rocky limestone slopes and hills, canyon bottoms and slopes, limestone rubble
Elevation 1500-1800 m (4900-5900 ft) 1200-1600 m (3900-5200 ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Coahuila)
[BONAP county map]
Discussion

Of conservation concern.

Physaria bellii is often found in shale and limestone soils of the Fountain/Ingleside, Lykins, Niobrara, and Pierre formations. It is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 628. FNA vol. 7, p. 650.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella mcvaughiana
Name authority G. A. Mulligan: Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. (1966) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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