Physaria bellii |
Physaria engelmannii |
|
---|---|---|
Bell's or Front Range twinpod, Bell's twinpod, Front Range twinpod |
Engelmann's bladderpod |
|
Habit | Perennials; caudex simple, (relatively large); densely (silvery) pubescent, trichomes (sessile, appressed), rays furcate, fused at base. | Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). |
Stems | simple from base, decumbent to nearly prostrate, 0.5–1.3 dm. |
few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm. |
Basal leaves | (strongly rosulate; shortly petiolate); blade broadly obovate, 1.5–7.5 (width 7.5–26 mm, base gradually tapering to petiole), margins shallowly dentate, (apex obtuse). |
blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent). |
Cauline leaves | blade oblanceolate to broadly obovate, 1–2.5 cm, margins entire. |
(proximal often petiolate, distal sessile or subsessile); blade oblanceolate to linear, 1–4 cm, margins entire. |
Racemes | dense. |
dense, (subumbellate). |
Flowers | sepals (pale yellow or yellow-green), narrowly lanceolate to narrowly deltate, 4–8 mm; petals yellow, broadly spatulate to obovate, 9–13 mm, (not clawed). |
sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse). |
Fruiting pedicels | (divaricate-ascending to widely spreading, slightly sigmoid to curved), 7–12 mm. |
(ascending), relatively short. |
Fruits | didymous, slightly flattened (contrary to replum) to uncompressed, 4–9 × 2–8 mm, (strongly coriaceous, apical and basal sinuses narrow, deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed; replum narrowly oblanceolate to narrowly linear-oblong, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style more than 3 mm. |
(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20 per ovary; style 3.5–5 mm. |
Seeds | compressed. |
flattened. |
2n | = 8. |
= 12, 24, 36. |
Physaria bellii |
Physaria engelmannii |
|
Phenology | Flowering Mar–Jun(-Jul). | Flowering Apr–May. |
Habitat | Dark shale, road cuts, ridge crests, washes | Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops |
Elevation | 1500-1800 m (4900-5900 ft) | 150-400 m (500-1300 ft) |
Distribution |
CO
|
OK; TX |
Discussion | Of conservation concern. Physaria bellii is often found in shale and limestone soils of the Fountain/Ingleside, Lykins, Niobrara, and Pierre formations. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 635. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella | |
Name authority | G. A. Mulligan: Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. (1966) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
Web links |