Physaria bellii |
Physaria arizonica |
|
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Bell's or Front Range twinpod, Bell's twinpod, Front Range twinpod |
Arizona bladderpod |
|
Habit | Perennials; caudex simple, (relatively large); densely (silvery) pubescent, trichomes (sessile, appressed), rays furcate, fused at base. | Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout). |
Stems | simple from base, decumbent to nearly prostrate, 0.5–1.3 dm. |
simple or few to several from base, erect, (unbranched, slender), 0.2–1(–1.5) dm. |
Basal leaves | (strongly rosulate; shortly petiolate); blade broadly obovate, 1.5–7.5 (width 7.5–26 mm, base gradually tapering to petiole), margins shallowly dentate, (apex obtuse). |
(densely tufted, not rosulate, reflexed in age); blade obovate to oblanceolate, 0.7–2(–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute). |
Cauline leaves | blade oblanceolate to broadly obovate, 1–2.5 cm, margins entire. |
similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5(–5.5) cm. |
Racemes | dense. |
dense, often subcorymbiform. |
Flowers | sepals (pale yellow or yellow-green), narrowly lanceolate to narrowly deltate, 4–8 mm; petals yellow, broadly spatulate to obovate, 9–13 mm, (not clawed). |
sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically); petals (spreading), oblanceolate to obovate, 5.5–8(–10) mm, (claw erect). |
Fruiting pedicels | (divaricate-ascending to widely spreading, slightly sigmoid to curved), 7–12 mm. |
(erect or divaricate-spreading, straight or slightly curved), (3–)5–10(–15) mm. |
Fruits | didymous, slightly flattened (contrary to replum) to uncompressed, 4–9 × 2–8 mm, (strongly coriaceous, apical and basal sinuses narrow, deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed; replum narrowly oblanceolate to narrowly linear-oblong, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style more than 3 mm. |
(sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm; valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth; ovules 4–10(–16) per ovary; style (0.5–)1–2(–4) mm (shorter than fruit). |
Seeds | compressed. |
flattened. |
2n | = 8. |
= 10. |
Physaria bellii |
Physaria arizonica |
|
Phenology | Flowering Mar–Jun(-Jul). | Flowering Apr–Jun. |
Habitat | Dark shale, road cuts, ridge crests, washes | Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine |
Elevation | 1500-1800 m (4900-5900 ft) | 1000-2200 m (3300-7200 ft) |
Distribution |
CO
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AZ; UT
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Discussion | Of conservation concern. Physaria bellii is often found in shale and limestone soils of the Fountain/Ingleside, Lykins, Niobrara, and Pierre formations. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 627. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella arizonica, Lesquerella arizonica var. nudicaulis, P. arizonica var. andrusensis | |
Name authority | G. A. Mulligan: Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. (1966) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
Web links |