Physaria bellii |
Physaria arenosa |
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Bell's or Front Range twinpod, Bell's twinpod, Front Range twinpod |
Great Plains bladderpod |
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Habit | Perennials; caudex simple, (relatively large); densely (silvery) pubescent, trichomes (sessile, appressed), rays furcate, fused at base. | Perennials or, rarely, annuals; caudex simple or branched; ± densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (usually spreading), distinct or slightly fused at base, furcate or bifurcate, (tuberculate). | ||||
Stems | simple from base, decumbent to nearly prostrate, 0.5–1.3 dm. |
simple or few from base, prostrate or straggling to erect, (sometimes purplish, usually unbranched), (0.5–)1–2(–3) dm. |
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Basal leaves | (strongly rosulate; shortly petiolate); blade broadly obovate, 1.5–7.5 (width 7.5–26 mm, base gradually tapering to petiole), margins shallowly dentate, (apex obtuse). |
blade oblanceolate, 1.5–5(–7) cm, margins entire or shallowly dentate, (flat). |
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Cauline leaves | blade oblanceolate to broadly obovate, 1–2.5 cm, margins entire. |
blade elliptic to linear, (0.5–)1–2.5(–3) cm, margins usually entire. |
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Racemes | dense. |
(secund), dense, (elongated in fruit). |
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Flowers | sepals (pale yellow or yellow-green), narrowly lanceolate to narrowly deltate, 4–8 mm; petals yellow, broadly spatulate to obovate, 9–13 mm, (not clawed). |
sepals elliptic or oblong, 4–6(–7) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (often red or lavender when dried), obovate, 6–8.5(–9.5) mm, (narrowing to broad claw). |
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Fruiting pedicels | (divaricate-ascending to widely spreading, slightly sigmoid to curved), 7–12 mm. |
(usually sharply recurved, sometimes divaricate-spreading or nearly horizontal), 5–15(–20) mm, (stout). |
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Fruits | didymous, slightly flattened (contrary to replum) to uncompressed, 4–9 × 2–8 mm, (strongly coriaceous, apical and basal sinuses narrow, deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed; replum narrowly oblanceolate to narrowly linear-oblong, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style more than 3 mm. |
subglobose, obovoid, or broadly ellipsoid, slightly inflated, (3.5–)4–5.5(–6.5) mm; valves densely pubescent outside, trichomes spreading or closely appressed, rarely sparsely pubescent inside; ovules (4–)8(–10) per ovary; style (slender), 3–5.5(–6.5) mm. |
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Seeds | compressed. |
slightly flattened. |
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2n | = 8. |
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Physaria bellii |
Physaria arenosa |
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Phenology | Flowering Mar–Jun(-Jul). | |||||
Habitat | Dark shale, road cuts, ridge crests, washes | |||||
Elevation | 1500-1800 m (4900-5900 ft) | |||||
Distribution |
CO
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CO; MT; ND; NE; SD; WY; AB; MB; SK
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Discussion | Of conservation concern. Physaria bellii is often found in shale and limestone soils of the Fountain/Ingleside, Lykins, Niobrara, and Pierre formations. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 626. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria arenosa, Lesquerella arenosa, Lesquerella argentea var. arenosa, Lesquerella ludoviciana var. arenosa | |||||
Name authority | G. A. Mulligan: Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. (1966) | (Richardson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | ||||
Web links |