Physaria bellii |
Physaria arctica |
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Bell's or Front Range twinpod, Bell's twinpod, Front Range twinpod |
arctic bladderpod |
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Habit | Perennials; caudex simple, (relatively large); densely (silvery) pubescent, trichomes (sessile, appressed), rays furcate, fused at base. | Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth). |
Stems | simple from base, decumbent to nearly prostrate, 0.5–1.3 dm. |
simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm. |
Basal leaves | (strongly rosulate; shortly petiolate); blade broadly obovate, 1.5–7.5 (width 7.5–26 mm, base gradually tapering to petiole), margins shallowly dentate, (apex obtuse). |
(usually ± rosulate); blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire. |
Cauline leaves | blade oblanceolate to broadly obovate, 1–2.5 cm, margins entire. |
(sessile or shortly petiolate); blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire. |
Racemes | dense. |
loose. |
Flowers | sepals (pale yellow or yellow-green), narrowly lanceolate to narrowly deltate, 4–8 mm; petals yellow, broadly spatulate to obovate, 9–13 mm, (not clawed). |
sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw). |
Fruiting pedicels | (divaricate-ascending to widely spreading, slightly sigmoid to curved), 7–12 mm. |
(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout). |
Fruits | didymous, slightly flattened (contrary to replum) to uncompressed, 4–9 × 2–8 mm, (strongly coriaceous, apical and basal sinuses narrow, deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed; replum narrowly oblanceolate to narrowly linear-oblong, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style more than 3 mm. |
subglobose to ellipsoid, uncompressed, 4–6(–9) mm; valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile; replum as wide as or wider than fruit; ovules (8–)10–14(–16) per ovary; style 1–2.5(–4) mm. |
Seeds | compressed. |
plump. |
2n | = 8. |
= 60. |
Physaria bellii |
Physaria arctica |
|
Phenology | Flowering Mar–Jun(-Jul). | Flowering May–Aug. |
Habitat | Dark shale, road cuts, ridge crests, washes | Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance |
Elevation | 1500-1800 m (4900-5900 ft) | 0-1800 m (0-5900 ft) |
Distribution |
CO
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AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia) |
Discussion | Of conservation concern. Physaria bellii is often found in shale and limestone soils of the Fountain/Ingleside, Lykins, Niobrara, and Pierre formations. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 628. | FNA vol. 7, p. 626. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa | |
Name authority | G. A. Mulligan: Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. (1966) | (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
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