Physaria arizonica |
Physaria pinetorum |
|
---|---|---|
Arizona bladderpod |
White Mountain bladderpod |
|
Habit | Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout). | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 6–8-rayed, rays furcate or bifurcate, (tuberculate, less so on outer layers). |
Stems | simple or few to several from base, erect, (unbranched, slender), 0.2–1(–1.5) dm. |
simple or few from base, ascending to erect, (0.5–)1–2(–3.5) dm. |
Basal leaves | (densely tufted, not rosulate, reflexed in age); blade obovate to oblanceolate, 0.7–2(–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute). |
(petiole tapering to blade); blade rhombic to elliptic and irregularly angular, sometimes spatulate to oblanceolate, 1.5–7.5(–10) cm, margins entire. |
Cauline leaves | similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5(–5.5) cm. |
(not or loosely overlapping, petiolate or distal sessile); blade spatulate to oblanceolate, 1–4 cm, margins entire. |
Racemes | dense, often subcorymbiform. |
crowded, elongated. |
Flowers | sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically); petals (spreading), oblanceolate to obovate, 5.5–8(–10) mm, (claw erect). |
sepals ovate, oblong, or elliptic 4–7.5 mm, (median pair thickened apically, cucullate); petals spatulate or broadly cuneate, 6–13 mm, (claw slightly expanded at base). |
Fruiting pedicels | (erect or divaricate-spreading, straight or slightly curved), (3–)5–10(–15) mm. |
(ascending, curved or sigmoid), 6–12(–20) mm. |
Fruits | (sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm; valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth; ovules 4–10(–16) per ovary; style (0.5–)1–2(–4) mm (shorter than fruit). |
(substipitate), globose or obovoid to ellipsoid, sometimes slightly obcompressed, 4–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–24 per ovary; style (2–)4–7 mm. |
Seeds | flattened. |
flattened. |
2n | = 10. |
= 10. |
Physaria arizonica |
Physaria pinetorum |
|
Phenology | Flowering Apr–Jun. | Flowering Apr–Jul. |
Habitat | Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine | Scrub oak, pinyon-juniper woodland, open ponderosa pine forests, these sometimes mixed with Douglas fir, white pine, white fir, Engelmann spruce, or Gambel oak, on limestone-derived or otherwise basic soils, often in rock crevices |
Elevation | 1000-2200 m (3300-7200 ft) | 1400-2900 (-3400) m (4600-9500 (-11200) ft) |
Distribution |
AZ; UT
|
AZ; NM
|
Discussion | The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria pinetorum with reduced forms are found at high elevations; in disturbed, moist soils plants can become quite large, as in the Manzano Mountains. Densely cespitose plants with crowded racemes not exceeding the basal leaves are found at the crest (3200–3400 m) of the Sandia Mountains, New Mexico. These probably represent an undescribed taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 627. | FNA vol. 7, p. 658. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella arizonica, Lesquerella arizonica var. nudicaulis, P. arizonica var. andrusensis | Lesquerella pinetorum |
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
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