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threadleaf bladderpod

Snake Range bladderpod

Habit Annuals; with a fine taproot; ± densely pubescent, trichomes several-rayed, rays distinct or fused at base, bifurcate, (prominently tuberculate throughout). Perennials; caudex simple; densely pubescent, trichomes 5–7-rayed, rays bifurcate or trifurcate.
Stems

simple or few to several from base, erect, (sometimes branched), to 4 dm.

several from base, erect or outer usually decumbent toward base, (from below a terminal tuft of mostly erect leaves, unbranched), 1–2 dm.

Basal leaves

blade elliptic to rhombic, 3–8 cm, (base narrowing gradually to petiole), margins entire, repand, coarsely toothed, or pinnatifid.

blade ovate to elliptic, 2–4 cm, (base gradually narrowed to petiole), margins entire, (surfaces densely pubescent with trichome layers).

Cauline leaves

(proximal often shortly petiolate, distal sessile);

blade linear or narrowly obovate, 1.5–6(–10) cm, margins entire, repand, or shallowly toothed.

(remote, proximal shortly petiolate);

blade spatulate to oblanceolate, similar to basal, (base often cuneate), margins entire.

Racemes

usually loose.

elongated.

Flowers

sepals elliptic or ovate, 4–6 mm, (lateral pair usually subsaccate);

petals obovate to obdeltate, 6–10 mm, (apex often emarginate).

sepals linear-oblong, 5–7 mm, (lateral pair subsaccate);

petals (erect), lingulate, 8–10 mm, (claw barely differentiated from blade).

Fruiting pedicels

(usually divaricate, sometimes horizontal, straight or slightly curved), 8–20 mm.

(recurved), 7–10 mm.

Fruits

not didymous, ± globose, slightly inflated, 4–6 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4 per ovary;

style 2–3.5 mm; (stigma expanded).

(usually pendent, sessile), subglobose, slightly flattened (angustiseptate), 4–5 mm;

valves densely pubescent, trichomes somewhat spreading;

ovules 8–12 per ovary;

style 4–5 mm.

Seeds

flattened, (margined).

plump, (mucilaginous when wetted).

2n

= 10.

Physaria angustifolia

Physaria pendula

Phenology Flowering Apr(-May). Flowering May–Jun.
Habitat Shallow limestone-derived soils, sometimes spreading to disturbed sites Limestone gravel and cobbles, typically with junipers
Elevation 90-300 m (300-1000 ft) 1700-2100 m (5600-6900 ft)
Distribution
from FNA
OK; TX
[BONAP county map]
from FNA
NV
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 625. FNA vol. 7, p. 657.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria angustifolia, Lesquerella angustifolia, Lesquerella longifolia Lesquerella pendula
Name authority (Nuttall ex Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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