Physaria angustifolia |
Physaria grahamii |
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threadleaf bladderpod |
Graham's twinpod |
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Habit | Annuals; with a fine taproot; ± densely pubescent, trichomes several-rayed, rays distinct or fused at base, bifurcate, (prominently tuberculate throughout). | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. |
Stems | simple or few to several from base, erect, (sometimes branched), to 4 dm. |
several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
Basal leaves | blade elliptic to rhombic, 3–8 cm, (base narrowing gradually to petiole), margins entire, repand, coarsely toothed, or pinnatifid. |
(outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
Cauline leaves | (proximal often shortly petiolate, distal sessile); blade linear or narrowly obovate, 1.5–6(–10) cm, margins entire, repand, or shallowly toothed. |
similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
Racemes | usually loose. |
loose, (elongated). |
Flowers | sepals elliptic or ovate, 4–6 mm, (lateral pair usually subsaccate); petals obovate to obdeltate, 6–10 mm, (apex often emarginate). |
sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
Fruiting pedicels | (usually divaricate, sometimes horizontal, straight or slightly curved), 8–20 mm. |
(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
Fruits | not didymous, ± globose, slightly inflated, 4–6 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4 per ovary; style 2–3.5 mm; (stigma expanded). |
didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
Seeds | flattened, (margined). |
plump, (suborbicular). |
2n | = 10. |
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Physaria angustifolia |
Physaria grahamii |
|
Phenology | Flowering Apr(-May). | Flowering May–Jun. |
Habitat | Shallow limestone-derived soils, sometimes spreading to disturbed sites | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay |
Elevation | 90-300 m (300-1000 ft) | 2100-2900 m (6900-9500 ft) |
Distribution |
OK; TX |
UT |
Discussion | Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 625. | FNA vol. 7, p. 642. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Vesicaria angustifolia, Lesquerella angustifolia, Lesquerella longifolia | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda |
Name authority | (Nuttall ex Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) |
Web links |