Physaria alpina |
Physaria spatulata |
|
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Avery Peak or alpine twinpod, Avery Peak twinpod |
alpine bladderpod, spatula-leaf bladderpod |
|
Habit | Perennials; (with a long taproot), caudex usually buried, simple, (enlarged, covered with marcescent leaf bases, crown rosulate and horizontal to somewhat ascending, forming a dense crown at apex of caudex); (silvery) pubescent throughout, trichomes (sessile or stipitate), 5–8-rayed, rays furcate or bifurcate, (rounded to umbonate, strongly tuberculate, less so or smooth over center). | Perennials; caudex simple, (relatively small); sparsely to moderately pubescent, trichomes 4- or 5-rayed, rays furcate or bifurcate, not fused, (tuberculate). |
Stems | few from base, decumbent, (arising laterally proximal to current season’s leaves), 0.3–0.8 dm. |
simple from base, erect to decumbent, (well-exserted beyond basal leaves, loosely spreading), 0.3–1.2 dm. |
Basal leaves | (petiole slender); blade broadly obovate, or deltate to ovate or narrower, 1.5–3.5 cm, (base abruptly to gradually narrowed to petiole), margins entire or obscurely few-toothed, (apex usually obtuse, nearly acute in narrower leaves). |
(erect to prostrate, petiole distinct from blade); blade (inner) spatulate to oblanceolate, or (outer) oblanceolate or orbicular, 1.5–4 cm, margins entire (rarely folded). |
Cauline leaves | (2–5 per stem); blade oblanceolate to spatulate, similar to basal, margins entire, (apex acute). |
blade spatulate, distinctly different from basal. |
Racemes | loose, (3–6-flowered). |
moderately dense, (6–20-flowered). |
Flowers | sepals narrowly oblong to linear, 7–9 mm; petals (erect), spatulate, 10–12(–15) mm. |
sepals (pale yellow), elliptic, 3.5–5 mm; petals lingulate, 6–9 mm. |
Fruiting pedicels | (widely spreading to ascending, slightly curved or straight), 7–11 mm. |
(strongly sigmoid), 10–20 mm (2 times longer than fruits). |
Fruits | (usually purplish in age), didymous, irregular and somewhat angular, not highly inflated, 4–11 × 10–13 mm, (coriaceous, papery, shallowly grooved distally and on sides, tapered and narrowed toward replum, base obtuse to truncate, apex with broad sinus to nearly truncate); valves (retaining seeds after dehiscence), densely pubescent, not silvery; replum elliptic to obovate, as wide as or wider than fruit, base rounded, margins sparsely pubescent or glabrous, apex rounded (with funicles); ovules 4 per ovary; style 5–7 mm, (glabrous). |
lanceolate or orbicular, slightly inflated, (2.5–)3–6 mm, (apex usually strongly narrowed); valves pubescent, trichomes sparse and closely appressed to surface; ovules 4–8 per ovary; style 2.5–6 mm (usually ± equal in length to mature fruit). |
Seeds | flattened. |
plump. |
Physaria alpina |
Physaria spatulata |
|
Phenology | Flowering Jun–Jul. | Flowering May-early Jul. |
Habitat | Whitish or red substrates from limestone or dolomite, ridge crests, rocky alpine tundra and open areas | Grasslands, subalpine meadows, sagebrush, scattered pines, fellfields, calcareous (sometimes alkaline) substrates |
Elevation | 3500-4000 m (11500-13100 ft) | 900-2900 m (3000-9500 ft) |
Distribution |
CO
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MT; ND; NE; SD; WY; AB; SK
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Discussion | Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 663. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella spatulata, Lesquerella alpina var. spatulata, Lesquerella nodosa, P. reediana subsp. spatulata, P. reediana var. spatulata | |
Name authority | Rollins: Brittonia 33: 339. (1981) | (Rydberg) Grady & O’Kane: Novon 17: 190. (2007) |
Web links |