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Avery Peak or alpine twinpod, Avery Peak twinpod

Mcvaugh's bladderpod

Habit Perennials; (with a long taproot), caudex usually buried, simple, (enlarged, covered with marcescent leaf bases, crown rosulate and horizontal to somewhat ascending, forming a dense crown at apex of caudex); (silvery) pubescent throughout, trichomes (sessile or stipitate), 5–8-rayed, rays furcate or bifurcate, (rounded to umbonate, strongly tuberculate, less so or smooth over center). Perennials; caudex simple or branched, (sometimes enlarged); densely pubescent, trichomes (sessile), several-rayed, rays fused (webbed) most of their length, (umbonate, peltate, tuberculate throughout).
Stems

few from base, decumbent, (arising laterally proximal to current season’s leaves), 0.3–0.8 dm.

few to several from base, erect or outer ones decumbent, 0.5–4 dm.

Basal leaves

(petiole slender);

blade broadly obovate, or deltate to ovate or narrower, 1.5–3.5 cm, (base abruptly to gradually narrowed to petiole), margins entire or obscurely few-toothed, (apex usually obtuse, nearly acute in narrower leaves).

(long-petiolate);

blade elliptic to obovate or rhombic, 2–6(–9) cm, margins entire.

Cauline leaves

(2–5 per stem);

blade oblanceolate to spatulate, similar to basal, margins entire, (apex acute).

(sessile or shortly petiolate);

blade oblanceolate to spatulate, 1–3 cm, (proximal broader), margins entire.

Racemes

loose, (3–6-flowered).

dense, (relatively short).

Flowers

sepals narrowly oblong to linear, 7–9 mm;

petals (erect), spatulate, 10–12(–15) mm.

sepals elliptic or narrowly oblong, 4–5.4 mm, (tapered to apex);

petals (white, base and claw yellow, conspicuously purple-veined), usually broadly obovate or rhombic, 6–10 mm, (± equal to blade, tapering to slender claw).

Fruiting pedicels

(widely spreading to ascending, slightly curved or straight), 7–11 mm.

(erect to spreading, ascending, or (proximal) horizontal, straight to slightly curved, sometimes loosely sigmoid), 6–12(–20) mm.

Fruits

(usually purplish in age), didymous, irregular and somewhat angular, not highly inflated, 4–11 × 10–13 mm, (coriaceous, papery, shallowly grooved distally and on sides, tapered and narrowed toward replum, base obtuse to truncate, apex with broad sinus to nearly truncate);

valves (retaining seeds after dehiscence), densely pubescent, not silvery;

replum elliptic to obovate, as wide as or wider than fruit, base rounded, margins sparsely pubescent or glabrous, apex rounded (with funicles);

ovules 4 per ovary;

style 5–7 mm, (glabrous).

(sessile or substipitate, often reddish magenta), usually ovoid to subglobose, inflated, 4–6(–7) mm;

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

septum perforate;

ovules 8–12 per ovary;

style 1.5–4 mm.

Seeds

flattened.

somewhat flattened.

2n

= 12.

Physaria alpina

Physaria mcvaughiana

Phenology Flowering Jun–Jul. Flowering mid Mar–Apr(-Aug).
Habitat Whitish or red substrates from limestone or dolomite, ridge crests, rocky alpine tundra and open areas Stream bed gravels, rocky limestone slopes and hills, canyon bottoms and slopes, limestone rubble
Elevation 3500-4000 m (11500-13100 ft) 1200-1600 m (3900-5200 ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Coahuila)
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 624. FNA vol. 7, p. 650.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella mcvaughiana
Name authority Rollins: Brittonia 33: 339. (1981) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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