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Avery Peak or alpine twinpod, Avery Peak twinpod

globe bladderpod

Habit Perennials; (with a long taproot), caudex usually buried, simple, (enlarged, covered with marcescent leaf bases, crown rosulate and horizontal to somewhat ascending, forming a dense crown at apex of caudex); (silvery) pubescent throughout, trichomes (sessile or stipitate), 5–8-rayed, rays furcate or bifurcate, (rounded to umbonate, strongly tuberculate, less so or smooth over center). Biennials or perennials; caudex branched, (± woody); densely pubescent, trichomes (sessile), 3–6-rayed, rays distinct and simple or furcate, (in 2 layers, lower layer umbonate, smooth to finely tuberculate, some often with a U-shaped notch).
Stems

few from base, decumbent, (arising laterally proximal to current season’s leaves), 0.3–0.8 dm.

several from base, erect, (arising among leaves of an elongated main axis), to 5 dm (± equal).

Basal leaves

(petiole slender);

blade broadly obovate, or deltate to ovate or narrower, 1.5–3.5 cm, (base abruptly to gradually narrowed to petiole), margins entire or obscurely few-toothed, (apex usually obtuse, nearly acute in narrower leaves).

(shortly petiolate);

blade obovate to oblanceolate, (1.5–)2.5–5(–6) cm, margins entire, sinuate to shallowly toothed, or pinnatifid.

Cauline leaves

(2–5 per stem);

blade oblanceolate to spatulate, similar to basal, margins entire, (apex acute).

(sessile or shortly petiolate);

blade oblanceolate to oblong, 1.3–3(–4) cm, (base cuneate), margins entire or repand to dentate.

Racemes

loose, (3–6-flowered).

dense.

Flowers

sepals narrowly oblong to linear, 7–9 mm;

petals (erect), spatulate, 10–12(–15) mm.

sepals elliptic or obovate, 2.6–4.1 mm, (median pair thickened apically, cucullate);

petals (bright yellow), obovate, 3.5–6.5(–7.5) mm, (margins sinuate).

Fruiting pedicels

(widely spreading to ascending, slightly curved or straight), 7–11 mm.

(usually spreading horizontally, straight), 7–14(–21) mm.

Fruits

(usually purplish in age), didymous, irregular and somewhat angular, not highly inflated, 4–11 × 10–13 mm, (coriaceous, papery, shallowly grooved distally and on sides, tapered and narrowed toward replum, base obtuse to truncate, apex with broad sinus to nearly truncate);

valves (retaining seeds after dehiscence), densely pubescent, not silvery;

replum elliptic to obovate, as wide as or wider than fruit, base rounded, margins sparsely pubescent or glabrous, apex rounded (with funicles);

ovules 4 per ovary;

style 5–7 mm, (glabrous).

(sessile or substipitate);

globose, often slightly compressed apically, (1–)2–3 mm;

valves sparsely pubescent, sometimes pubescent inside, trichomes spreading, 3–5-rayed;

ovules 4 per ovary;

style 2–3.5(–4) mm.

Seeds

flattened.

flattened or plump, (often outer surface hemispherical, inner surface flattened, or both surfaces rounded).

2n

= 14.

Physaria alpina

Physaria globosa

Phenology Flowering Jun–Jul. Flowering Mar–May.
Habitat Whitish or red substrates from limestone or dolomite, ridge crests, rocky alpine tundra and open areas Open rocky areas, shale at cliff bases, open talus, ledges, open cedar glades
Elevation 3500-4000 m (11500-13100 ft) 100-300 m (300-1000 ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
IN; KY; TN
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria globosa is possibly introduced in Indiana. A report for Ohio was based on a collection by “Jones,” but that specimen cannot be located.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 624. FNA vol. 7, p. 639.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria globosa, Alyssum globosum, Alyssum shortii, Lesquerella globosa
Name authority Rollins: Brittonia 33: 339. (1981) (Desvaux) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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