Physaria alpina |
Physaria arizonica |
|
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Avery Peak or alpine twinpod, Avery Peak twinpod |
Arizona bladderpod |
|
Habit | Perennials; (with a long taproot), caudex usually buried, simple, (enlarged, covered with marcescent leaf bases, crown rosulate and horizontal to somewhat ascending, forming a dense crown at apex of caudex); (silvery) pubescent throughout, trichomes (sessile or stipitate), 5–8-rayed, rays furcate or bifurcate, (rounded to umbonate, strongly tuberculate, less so or smooth over center). | Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout). |
Stems | few from base, decumbent, (arising laterally proximal to current season’s leaves), 0.3–0.8 dm. |
simple or few to several from base, erect, (unbranched, slender), 0.2–1(–1.5) dm. |
Basal leaves | (petiole slender); blade broadly obovate, or deltate to ovate or narrower, 1.5–3.5 cm, (base abruptly to gradually narrowed to petiole), margins entire or obscurely few-toothed, (apex usually obtuse, nearly acute in narrower leaves). |
(densely tufted, not rosulate, reflexed in age); blade obovate to oblanceolate, 0.7–2(–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute). |
Cauline leaves | (2–5 per stem); blade oblanceolate to spatulate, similar to basal, margins entire, (apex acute). |
similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5(–5.5) cm. |
Racemes | loose, (3–6-flowered). |
dense, often subcorymbiform. |
Flowers | sepals narrowly oblong to linear, 7–9 mm; petals (erect), spatulate, 10–12(–15) mm. |
sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically); petals (spreading), oblanceolate to obovate, 5.5–8(–10) mm, (claw erect). |
Fruiting pedicels | (widely spreading to ascending, slightly curved or straight), 7–11 mm. |
(erect or divaricate-spreading, straight or slightly curved), (3–)5–10(–15) mm. |
Fruits | (usually purplish in age), didymous, irregular and somewhat angular, not highly inflated, 4–11 × 10–13 mm, (coriaceous, papery, shallowly grooved distally and on sides, tapered and narrowed toward replum, base obtuse to truncate, apex with broad sinus to nearly truncate); valves (retaining seeds after dehiscence), densely pubescent, not silvery; replum elliptic to obovate, as wide as or wider than fruit, base rounded, margins sparsely pubescent or glabrous, apex rounded (with funicles); ovules 4 per ovary; style 5–7 mm, (glabrous). |
(sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm; valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth; ovules 4–10(–16) per ovary; style (0.5–)1–2(–4) mm (shorter than fruit). |
Seeds | flattened. |
flattened. |
2n | = 10. |
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Physaria alpina |
Physaria arizonica |
|
Phenology | Flowering Jun–Jul. | Flowering Apr–Jun. |
Habitat | Whitish or red substrates from limestone or dolomite, ridge crests, rocky alpine tundra and open areas | Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine |
Elevation | 3500-4000 m (11500-13100 ft) | 1000-2200 m (3300-7200 ft) |
Distribution |
CO
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AZ; UT
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Discussion | Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 627. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella arizonica, Lesquerella arizonica var. nudicaulis, P. arizonica var. andrusensis | |
Name authority | Rollins: Brittonia 33: 339. (1981) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
Web links |