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Physaria alpestris

alpine twin-pod, Washington bladder-pod, Washington twin-pod

straight bladderpod

Habit Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). Perennials; (loose, spreading); caudex simple or branched; densely pubescent, trichomes (subsessile), 4–6-rayed, rays furcate or bifurcate, (moderately tuberculate over arms, less so or smooth over center).
Stems

several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm.

few to several from base, ascending or prostrate, (arising laterally, also from within basal leaves, usually unbranched, rarely branched), 0.5–3(–6) dm.

Basal leaves

(petiole slender);

blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute).

blade narrowly oblanceolate or broadly elliptic, 1–7(–12) cm, margins entire or shallowly toothed, sometimes repand, (inner blades usually flattened in age, surfaces often gray-green, scabrous).

Cauline leaves

blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire.

(usually secund);

blade spatulate or obovate, 1–2.5(–4.5) cm, margins entire or shallowly toothed, (flat or involute).

Racemes

subcorymbose.

somewhat crowded (to moderately elongated in fruit, exceeding basal leaves).

Flowers

sepals oblong, 8–10 mm;

petals spatulate, 12–14 mm.

sepals broadly elliptic or oblong, 4–7.5(–9) mm, (median pair thickened apically, cucullate);

petals cuneate or obovate, 7–10(–16) mm, (tapering gradually to broad claw).

Fruiting pedicels

(divaricate, straight), 5–10 mm.

(often divaricate-spreading and straight, or horizontal and loosely sigmoid, sometimes slightly recurved), 5–15 mm.

Fruits

didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow);

valves (retaining seeds after dehiscence), evenly pubescent;

replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate;

ovules 8–10 per ovary;

style 5–7 mm.

subglobose to ovoid or ellipsoid, sometimes compressed, (4–)5–7(–9) mm;

valves sparsely pubescent, trichomes appressed or erect, sometimes sparsely pubescent inside;

ovules (8–)12–16(–20) per ovary;

style 2–7 mm.

Seeds

flattened, (2–3 mm).

somewhat flattened.

2n

= 48–52, 52, 64, 67–70.

= 10 + 2, 18, 20, ± 40.

Physaria alpestris

Physaria rectipes

Phenology Flowering May–Jun. Flowering May–Jul.
Habitat Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities Sandy soils, limey knolls, rocky hills, clay hillsides, dry ridges, weathered rocks, gravelly outwashes, stony slopes, pinyon-juniper woodlands
Elevation (700-)1300-2400 m ((2300-)4300-7900 ft) 1500-2600 m (4900-8500 ft)
Distribution
from FNA
WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

As here circumscribed, Physaria rectipes remains heterogeneous and may represent more than one taxon, even after the recent removal of P. pulvinata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 624. FNA vol. 7, p. 660.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella alpestris Lesquerella rectipes
Name authority Suksdorf: W. Amer. Sci. 15: 58. (1906) (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002)
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