Physaria alpestris |
Physaria pruinosa |
|
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
frosty bladderpod, Pagosa bladderpod |
|
Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Perennials; caudex simple or branched, (covered with persistent leaf bases); densely pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout). |
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
simple or several from base, decumbent or erect, (unbranched), to 2 dm. |
Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
(petiole sharply differentiated from blade, slender); blade suborbicular or obovate to rhombic, 4–8 cm, (base abruptly narrowed to petiole), margins entire, sinuate, or shallowly dentate, (abaxial surface densely pubescent, adaxial lightly pubescent). |
Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
(proximal petiolate, distal sessile); blade obovate to rhombic, 0.8–2.3 cm, margins entire or shallowly toothed. |
Racemes | subcorymbose. |
dense, (somewhat elongated in fruit). |
Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals elliptic or oblong, ca. 6 mm, (lateral pair not saccate or subsaccate, cucullate, median pair thickened apically, cucullate); petals spatulate, ca. 9 mm, (claw expanded at base). |
Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(horizontal to ascending, sigmoid or slightly curved), 8–11 mm, (stout). |
Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
(sessile or substipitate, often becoming copper-red in age), subglobose or ellipsoid, inflated, 6–9 mm, (firm, glossy); valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8(–12) per ovary; style 3.5–7 mm. |
Seeds | flattened, (2–3 mm). |
somewhat flattened. |
2n | = 48–52, 52, 64, 67–70. |
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Physaria alpestris |
Physaria pruinosa |
|
Phenology | Flowering May–Jun. | Flowering May–Jun(-Aug). |
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | Mancos slate or shale, meadows, gentle slopes, edges of ponderosa pine stands |
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | 2100-2600 m (6900-8500 ft) |
Distribution |
WA
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CO; NM |
Discussion | Of conservation concern. The one New Mexico population is near the border with Colorado, in Rio Arriba County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 658. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella alpestris | Lesquerella pruinosa |
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | (Greene) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
Web links |