Physaria alpestris |
Physaria nelsonii |
|
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
Nelson's bladderpod |
|
Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Perennials; (diminutive); caudex branched, (densely cespitose, mound-forming); densely pubescent, trichomes (flaring, giving a shaggy appearance), usually 4–5-rayed, rays furcate or bifurcate, (strongly tuberculate throughout). |
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
few to several from base, erect, (from basal tuft), 0.1–0.2(–0.3) dm, (not or just barely exceeding leaves). |
Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
blade spatulate to oblanceolate, 0.5–1.5 cm, (base gradually narrowed to petiole), margins entire. |
Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
(absent or few); similar to basal, blade linear. |
Racemes | subcorymbose. |
dense, (few-flowered). |
Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals (pale yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate); petals (bright yellow), lingulate, 6–8 mm. |
Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(loosely sigmoid), 3–5 mm. |
Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
lanceolate, compressed apically, 2.5–3(–4) mm; valves pubescent, trichomes spreading, (appearing fuzzy), rarely with trichomes inside; ovules 4–8 per ovary; style 2.5–4 mm. |
Seeds | flattened, (2–3 mm). |
plump, (oblong). |
2n | = 48–52, 52, 64, 67–70. |
|
Physaria alpestris |
Physaria nelsonii |
|
Phenology | Flowering May–Jun. | Flowering May-mid Jun. |
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | Limestone, windswept knolls and cliffs, nearly barren areas with other cushion-forming plants |
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | 1600-2300 m (5200-7500 ft) |
Distribution |
WA
|
UT; WY |
Discussion | Physaria nelsonii is morphologically similar to 73. P. pycnantha, which traditionally was included in a broader P. nelsonii. These allopatric species are distinguished by styles equal to or exceeding the length of fruit and fruiting stems overtopped by basal leaves (P. nelsonii) versus styles shorter than fruits and fruiting stems usually exserted beyond basal leaves (P. pycnantha). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 652. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella alpestris | Lesquerella condensata, Lesquerella alpina subsp. condensata, Lesquerella alpina var. condensata |
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | O’Kane & Al-Shehbaz: Novon 12: 326. (2002) |
Web links |