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Physaria alpestris

alpine twin-pod, Washington bladder-pod, Washington twin-pod

largefruit bladderpod

Habit Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). Perennials; caudex branched; densely pubescent, trichomes (sessile or short-stalked), 4–6-rayed, rays distinct, usually furcate, rarely bifurcate, (finely tuberculate throughout).
Stems

several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm.

few or several from base, prostrate to decumbent, (unbranched or branched), 0.5–1.5 dm.

Basal leaves

(petiole slender);

blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute).

blades orbicular to broadly obovate, 1.5–3 cm, margins usually entire, rarely remotely dentate.

Cauline leaves

blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire.

(sessile or shortly petiolate);

blade elliptic to oblanceolate, 1–1.5(–2.5) cm, margins entire, (apex obtuse).

Racemes

subcorymbose.

dense, (elongated in fruit).

Flowers

sepals oblong, 8–10 mm;

petals spatulate, 12–14 mm.

sepals ovate or oblong-elliptic, 5–5.5 mm, (lateral pair not saccate);

petals cuneate or broadly obovate, ca. 7 mm, (sometimes slightly narrowed to a broad claw, apex sometimes retuse).

Fruiting pedicels

(divaricate, straight), 5–10 mm.

(sharply recurved), 5–10 mm, (stout).

Fruits

didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow);

valves (retaining seeds after dehiscence), evenly pubescent;

replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate;

ovules 8–10 per ovary;

style 5–7 mm.

subglobose to broadly obovoid, strongly inflated (often slightly angustiseptate), 5–7 mm, (papery);

valves sparsely pubescent; (septum fenestrate, perforate, or obsolete);

ovules 4–8 per ovary;

style 2–3 mm.

Seeds

flattened, (2–3 mm).

somewhat flattened.

2n

= 48–52, 52, 64, 67–70.

Physaria alpestris

Physaria macrocarpa

Phenology Flowering May–Jun. Flowering May–Jun.
Habitat Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities Gypsum-clay hills and benches, naked clay flats and barren hills
Elevation (700-)1300-2400 m ((2300-)4300-7900 ft) 2000-2400 m (6600-7900 ft)
Distribution
from FNA
WA
[WildflowerSearch map]
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Of conservation concern.

Physaria macrocarpa is found in the Great Divide and Green River basins.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 624. FNA vol. 7, p. 650.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella alpestris Lesquerella macrocarpa
Name authority Suksdorf: W. Amer. Sci. 15: 58. (1906) (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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