Physaria alpestris |
Physaria ludoviciana |
|
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
foothill bladderpod, silver bladderpod |
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Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). |
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
(erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
Racemes | subcorymbose. |
compact, (elongated and loose in fruit, densely-flowered). |
Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(usually recurved), (5–)10–20(–25) mm. |
Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
Seeds | flattened, (2–3 mm). |
slightly flattened. |
2n | = 48–52, 52, 64, 67–70. |
= 10, 20, 30. |
Physaria alpestris |
Physaria ludoviciana |
|
Phenology | Flowering May–Jun. | Flowering Apr–Jul(-Aug). |
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands |
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | 0-1900 m (0-6200 ft) |
Distribution |
WA
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AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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Discussion | Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 649. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella alpestris | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana |
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |