Physaria alpestris |
Physaria kingii |
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
King bladderpod, King's bladder-pod |
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Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). | ||||||||||||||||||||||||
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm. |
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Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed. |
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Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
(proximal petiolate, distal sessile); blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire. |
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Racemes | subcorymbose. |
(usually not secund), dense, (sometimes elongated in fruit). |
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Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals lanceolate, 4–6(–7) mm; petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade). |
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Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm. |
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Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
(sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute); valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete); ovules 4–16 per ovary; style 1–9 mm. |
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Seeds | flattened, (2–3 mm). |
flattened (sometimes slightly). |
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2n | = 48–52, 52, 64, 67–70. |
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Physaria alpestris |
Physaria kingii |
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Phenology | Flowering May–Jun. | |||||||||||||||||||||||||
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | |||||||||||||||||||||||||
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | |||||||||||||||||||||||||
Distribution |
WA
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AZ; CA; ID; NV; OR; UT; nw Mexico
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Discussion | Subspecies 7 (7 in the flora). The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 645. | ||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||
Synonyms | Lesquerella alpestris | Vesicaria kingii, Lesquerella kingii | ||||||||||||||||||||||||
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||||||||||||||||||||||
Web links |