Physaria alpestris |
Physaria integrifolia |
|
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
Snake River or creeping twinpod, Snake River twinpod |
|
Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Perennials; caudex usually branched, (rhizomelike, cespitose); densely (silvery) pubescent, trichomes (often stalked, appressed), several-rayed, rays furcate or bifurcate, (umbonate, strongly tuberculate throughout). |
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
several from base, ± erect, exceeding basal rosette by ± 0.5 dm. |
Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
(forming a strong rosette; long-petiolate); blade oblanceolate to ovate or orbicular, (1.5–)2–4(–8) cm, (base usually abruptly tapering to petiole), margins entire. |
Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
blade oblanceolate, 1–2 cm, margin entire, (apex acute). |
Racemes | subcorymbose. |
congested, (greatly exceeding leaves). |
Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals often keeled, 6–8 mm; petals spatulate, 8–10 mm. |
Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(spreading, straight or slightly curved), 7–11 mm. |
Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
didymous, highly inflated, 8–22 × 10–25 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), densely pubescent, trichomes appressed; replum linear to oblong, as wide as or wider than fruit; ovules 8 per ovary; style 7–9 mm. |
Seeds | flattened, (2–3 mm). |
flattened. |
2n | = 48–52, 52, 64, 67–70. |
= 16. |
Physaria alpestris |
Physaria integrifolia |
|
Phenology | Flowering May–Jun. | Flowering (May-)Jun–Jul(-Aug). |
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | Calcareous hills and slopes, shale-limestone cliffs, bare steep slopes, red clay banks, shale |
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | 1900-2700 m (6200-8900 ft) |
Distribution |
WA
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ID; MT; WY |
Discussion | Physaria integrifolia has traditionally been recognized as a variety of P. didymocarpa, but it is morphologically and ecologically quite distinctive. Variety monticola (no combination has been made at subspecific rank) is not recognized here; it is considered another example, in the genus, of caudices elongating in response to shifting substrates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 644. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella alpestris | P. didymocarpa var. integrifolia, P. integrifolia var. monticola |
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | (Rollins) Lichvar: Madroño 31: 203. (1984) |
Web links |