Physaria alpestris |
Physaria gracilis |
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
spreading bladderpod |
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Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). | ||||
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. |
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Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). |
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Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
(proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. |
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Racemes | subcorymbose. |
loose, (elongated). |
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Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). |
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Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). |
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Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
(stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. |
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Seeds | flattened, (2–3 mm). |
slightly flattened. |
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2n | = 48–52, 52, 64, 67–70. |
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Physaria alpestris |
Physaria gracilis |
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Phenology | Flowering May–Jun. | |||||
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | |||||
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | |||||
Distribution |
WA
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AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 641. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella alpestris | Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis | ||||
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |