Physaria alpestris |
Physaria gooddingii |
|
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
Goodding's bladderpod |
|
Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Annuals or biennials; without caudex, (cespitose); densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (ascending or erect), simple or infrequently furcate near base, (long and slender, sometimes with U-shaped notch on one side, smooth or finely tuberculate). |
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
several from base, erect (and stout) or outer ones decumbent, (unbranched or branched, stiff and densely foliate, sterile leaf-bearing branches sometimes present), to 4 dm. |
Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
blade obovate or elliptic, to ca. 3 cm, margins sinuate or shallowly dentate. |
Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
(proximal usually shortly petiolate, distal sessile); blade obovate to broadly elliptic, 1–3 cm, margins sinuate or shallowly toothed. |
Racemes | subcorymbose. |
dense, compact, (elongated in fruit). |
Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals elliptic or narrowly elliptic or oblong, (3.8–)4.5–5.5 mm, (lateral pair cucullate, very convex, median pair tapering to base, thickened apically, cucullate, often slightly keeled); petals cuneate, 6.5–8 mm, (slightly expanded at base, margins sinuate, apex retuse or entire). |
Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(recurved, curved or sigmoid), somewhat expanded apically. |
Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
(sessile or substipitate), oblong or broadly elliptic, compressed (latiseptate), 5–8 mm; valves pubescent, trichomes spreading, sparsely pubescent inside; ovules 4–6 per ovary; style 3–5 mm. |
Seeds | flattened, (2–3 mm). |
flattened. |
2n | = 48–52, 52, 64, 67–70. |
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Physaria alpestris |
Physaria gooddingii |
|
Phenology | Flowering May–Jun. | Flowering Jun–Sep. |
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | Mountainous areas, open areas in pinyon-juniper and ponderosa pine forests |
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | 1800-2300 m (5900-7500 ft) |
Distribution |
WA
|
AZ; NM |
Discussion | Physaria gooddingii (found in the mountains of Catron, Sierra, and western Socorro counties, New Mexico, and in Greenlee County, Arizona) is similar to 9. P. aurea (found farther east), but differs in having trichomes with ascending or erect rays (rather than appressed) and fruits that are strongly latiseptate (rather than not, or very little, compressed), a state that is infrequent in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 639. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella alpestris | Lesquerella gooddingii |
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) |
Web links |