Physaria alpestris |
Physaria fendleri |
|
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
Fendler bladderpod, Fendler's bladderpod |
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Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Perennials; caudex branched, (sometimes woody at base); densely (silvery) pubescent, trichomes (sessile or short-stalked), several-rayed, rays not furcate, fused (webbed) ca. 1/2 their length, (tuberculate throughout or tubercles scarce or absent over center). |
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
several from base, erect or laterally decumbent, (usually unbranched), (0.3–)0.5–2.5(–4) dm. |
Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
blade linear to somewhat elliptic, 1–4(–8) cm, margins entire or coarsely dentate. |
Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
(shortly petiolate); blade usually linear to narrowly oblanceolate, rarely elliptic to rhombic, 0.5–2.5 cm, (base narrowing to petiole), margins entire or remotely dentate (sometimes involute). |
Racemes | subcorymbose. |
loose to somewhat dense. |
Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals elliptic to oblong, 5–8 mm, (lateral pair not saccate, median pair often thickened apically, ± cucullate); petals (usually orange or orange-yellow at junction of blade and claw, sometimes also with orange guidelines), obdeltate to obovate, 8–12 mm, (claw relatively short). |
Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(divaricate-spreading to erect, usually straight or slightly curved, occasionally sigmoid), 8–20(–40) mm. |
Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
globose, broadly ellipsoid, or ovoid, not or slightly inflated, 5–8 mm, (firm, apex usually acute); valves (not retaining seeds after dehiscence, often reddish in age), glabrous throughout; replum as wide as or wider than fruit; ovules (12–)20–32(–40) per ovary; style (2–)3–6 mm. |
Seeds | flattened, (2–3 mm). |
flattened. |
2n | = 48–52, 52, 64, 67–70. |
= 12, 14, 24. |
Physaria alpestris |
Physaria fendleri |
|
Phenology | Flowering May–Jun. | Flowering Mar–May. |
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | Limestone outcrops, gypseous hills, gravels, sandy washes, rocky slopes, bluffs, shallow drainage areas, plains and desert shrub areas |
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | 100-2000 m (300-6600 ft) |
Distribution |
WA
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AZ; CO; KS; NM; OK; TX; UT; Mexico (Chihuahua, Coahuila, Nuevo León)
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Discussion | In dry areas, Physaria fendleri may flower following suitable rains at any time of the year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 636. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella alpestris | Vesicaria fendleri, Alyssum fendleri, Alyssum stenophyllum, Lesquerella fendleri, Lesquerella foliacea, Lesquerella praecox, Lesquerella stenophylla, Vesicaria stenophylla, Vesicaria stenophylla var. diffusa, Vesicaria stenophylla var. humilis, Vesicaria stenophylla var. procera |
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) |
Web links |