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Physaria alpestris

alpine twin-pod, Washington bladder-pod, Washington twin-pod

Engelmann's bladderpod

Habit Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate).
Stems

several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm.

few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm.

Basal leaves

(petiole slender);

blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute).

blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent).

Cauline leaves

blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire.

(proximal often petiolate, distal sessile or subsessile);

blade oblanceolate to linear, 1–4 cm, margins entire.

Racemes

subcorymbose.

dense, (subumbellate).

Flowers

sepals oblong, 8–10 mm;

petals spatulate, 12–14 mm.

sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate);

petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse).

Fruiting pedicels

(divaricate, straight), 5–10 mm.

(ascending), relatively short.

Fruits

didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow);

valves (retaining seeds after dehiscence), evenly pubescent;

replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate;

ovules 8–10 per ovary;

style 5–7 mm.

(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules (8–)12–20 per ovary;

style 3.5–5 mm.

Seeds

flattened, (2–3 mm).

flattened.

2n

= 48–52, 52, 64, 67–70.

= 12, 24, 36.

Physaria alpestris

Physaria engelmannii

Phenology Flowering May–Jun. Flowering Apr–May.
Habitat Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops
Elevation (700-)1300-2400 m ((2300-)4300-7900 ft) 150-400 m (500-1300 ft)
Distribution
from FNA
WA
[WildflowerSearch map]
[BONAP county map]
from FNA
OK; TX
[BONAP county map]
Source FNA vol. 7, p. 624. FNA vol. 7, p. 635.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella alpestris Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella
Name authority Suksdorf: W. Amer. Sci. 15: 58. (1906) (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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