Physaria alpestris |
Physaria didymocarpa |
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
common twinpod, double twinpod, twin bladderpod |
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Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | ||||||||
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
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Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
(forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
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Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
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Racemes | subcorymbose. |
congested, (elongated in fruit, greatly exceeding leaves). |
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Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
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Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(spreading, straight or slightly curved), 8–12 mm. |
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Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
(erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
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Seeds | flattened, (2–3 mm). |
flattened. |
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2n | = 48–52, 52, 64, 67–70. |
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Physaria alpestris |
Physaria didymocarpa |
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Phenology | Flowering May–Jun. | |||||||||
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | |||||||||
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | |||||||||
Distribution |
WA
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ID; MT; WA; WY; AB; BC
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Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 633. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lesquerella alpestris | Vesicaria didymocarpa, Coulterina didymocarpa | ||||||||
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | ||||||||
Web links |