Physaria alpestris |
Physaria arenosa |
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
Great Plains bladderpod |
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Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Perennials or, rarely, annuals; caudex simple or branched; ± densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (usually spreading), distinct or slightly fused at base, furcate or bifurcate, (tuberculate). | ||||
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
simple or few from base, prostrate or straggling to erect, (sometimes purplish, usually unbranched), (0.5–)1–2(–3) dm. |
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Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
blade oblanceolate, 1.5–5(–7) cm, margins entire or shallowly dentate, (flat). |
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Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
blade elliptic to linear, (0.5–)1–2.5(–3) cm, margins usually entire. |
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Racemes | subcorymbose. |
(secund), dense, (elongated in fruit). |
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Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
sepals elliptic or oblong, 4–6(–7) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (often red or lavender when dried), obovate, 6–8.5(–9.5) mm, (narrowing to broad claw). |
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Fruiting pedicels | (divaricate, straight), 5–10 mm. |
(usually sharply recurved, sometimes divaricate-spreading or nearly horizontal), 5–15(–20) mm, (stout). |
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Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
subglobose, obovoid, or broadly ellipsoid, slightly inflated, (3.5–)4–5.5(–6.5) mm; valves densely pubescent outside, trichomes spreading or closely appressed, rarely sparsely pubescent inside; ovules (4–)8(–10) per ovary; style (slender), 3–5.5(–6.5) mm. |
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Seeds | flattened, (2–3 mm). |
slightly flattened. |
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2n | = 48–52, 52, 64, 67–70. |
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Physaria alpestris |
Physaria arenosa |
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Phenology | Flowering May–Jun. | |||||
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | |||||
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | |||||
Distribution |
WA
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CO; MT; ND; NE; SD; WY; AB; MB; SK
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 626. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella alpestris | Vesicaria arenosa, Lesquerella arenosa, Lesquerella argentea var. arenosa, Lesquerella ludoviciana var. arenosa | ||||
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | (Richardson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | ||||
Web links |