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Physaria alpestris

alpine twin-pod, Washington bladder-pod, Washington twin-pod

Habit Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). Annuals, biennials, perennials, or subshrubs; eglandular.
Stems

several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm.

Basal leaves

(petiole slender);

blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute).

Cauline leaves

blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire.

petiolate, sessile, or subsessile;

blade base usually not auriculate (except Paysonia), margins entire, dentate, or sinuate.

Racemes

subcorymbose.

ebracteate, often elongated in fruit.

Flowers

sepals oblong, 8–10 mm;

petals spatulate, 12–14 mm.

actinomorphic;

sepals erect, spreading, ascending, or reflexed, lateral pair seldom saccate basally;

petals white, yellow, lavender, purple, violet, orange, or brown [pink], claw present, often distinct;

filaments unappendaged, not winged;

pollen (3 or) 4–11-colpate.

Fruiting pedicels

(divaricate, straight), 5–10 mm.

Fruits

didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow);

valves (retaining seeds after dehiscence), evenly pubescent;

replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate;

ovules 8–10 per ovary;

style 5–7 mm.

silicles or siliques, dehiscent, unsegmented, terete, latiseptate, or angustiseptate;

ovules 2–100 per ovary;

style usually distinct;

stigma entire or strongly 2-lobed.

Seeds

flattened, (2–3 mm).

biseriate, uniseriate, or aseriate;

cotyledons accumbent or incumbent.

Trichomes

usually short-stalked, subsessile, or sessile, sometimes long-stalked, stellate, scalelike, subdendritic, or forked, sometimes mixed with simple ones.

2n

= 48–52, 52, 64, 67–70.

Physaria alpestris

Brassicaceae tribe Physarieae

Phenology Flowering May–Jun.
Habitat Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities
Elevation (700-)1300-2400 m ((2300-)4300-7900 ft)
Distribution
from FNA
WA
[WildflowerSearch map]
[BONAP county map]
North America; Mexico; South America; Asia (ne Russia)
Discussion

Genera 7, species ca. 130 (7 genera, 105 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 624. FNA vol. 7, p. 604.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae
Sibling taxa
P. acutifolia, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
Synonyms Lesquerella alpestris
Name authority Suksdorf: W. Amer. Sci. 15: 58. (1906) B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 100. (1895)
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