Physaria alpestris |
Brassicaceae tribe Physarieae |
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alpine twin-pod, Washington bladder-pod, Washington twin-pod |
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Habit | Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). | Annuals, biennials, perennials, or subshrubs; eglandular. |
Stems | several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm. |
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Basal leaves | (petiole slender); blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute). |
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Cauline leaves | blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire. |
petiolate, sessile, or subsessile; blade base usually not auriculate (except Paysonia), margins entire, dentate, or sinuate. |
Racemes | subcorymbose. |
ebracteate, often elongated in fruit. |
Flowers | sepals oblong, 8–10 mm; petals spatulate, 12–14 mm. |
actinomorphic; sepals erect, spreading, ascending, or reflexed, lateral pair seldom saccate basally; petals white, yellow, lavender, purple, violet, orange, or brown [pink], claw present, often distinct; filaments unappendaged, not winged; pollen (3 or) 4–11-colpate. |
Fruiting pedicels | (divaricate, straight), 5–10 mm. |
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Fruits | didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow); valves (retaining seeds after dehiscence), evenly pubescent; replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate; ovules 8–10 per ovary; style 5–7 mm. |
silicles or siliques, dehiscent, unsegmented, terete, latiseptate, or angustiseptate; ovules 2–100 per ovary; style usually distinct; stigma entire or strongly 2-lobed. |
Seeds | flattened, (2–3 mm). |
biseriate, uniseriate, or aseriate; cotyledons accumbent or incumbent. |
Trichomes | usually short-stalked, subsessile, or sessile, sometimes long-stalked, stellate, scalelike, subdendritic, or forked, sometimes mixed with simple ones. |
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2n | = 48–52, 52, 64, 67–70. |
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Physaria alpestris |
Brassicaceae tribe Physarieae |
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Phenology | Flowering May–Jun. | |
Habitat | Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities | |
Elevation | (700-)1300-2400 m ((2300-)4300-7900 ft) | |
Distribution |
WA
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North America; Mexico; South America; Asia (ne Russia) |
Discussion | Genera 7, species ca. 130 (7 genera, 105 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 604. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Lesquerella alpestris | |
Name authority | Suksdorf: W. Amer. Sci. 15: 58. (1906) | B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 100. (1895) |
Web links |