Physaria acutifolia |
Physaria recurvata |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
gaslight bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Annuals or, sometimes, bi-ennials; with a fine taproot; sparsely pubescent, trichomes (sessile), 3–6-rayed, rays furcate, (tuberculate throughout). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
several from base, erect or decumbent and straggling, (branched distally, branches usually filiform), to 5 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blade obovate or rhombic to broadly elliptic, 1–4.5(–6.5) cm, margins entire or lyrate-pinnatifid. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(proximal petiolate, distal sessile); blade rhombic or obovate to elliptic, 0.5–2(–3) cm, margins entire or sinuate to remotely toothed. |
Racemes | loose, (elongated in fruit). |
loose. |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals elliptic or ovate, 2.5–5.5 mm, (median pair thickened apically, cucullate); petals (yellow to orange-yellow), obovate to cuneate, 4–7 mm, (apex sometimes retuse). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(recurved in age), 5–10(–15) mm, (slender). |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
globose or subglobose, not or slightly inflated, (2–)3–5(–7) mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (4–)8–16(–20) per ovary; style (1–5–)2–4.5 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
flattened. |
2n | = 10, 16, 24. |
= 10. |
Physaria acutifolia |
Physaria recurvata |
|
Phenology | Flowering May–Jun(-Jul). | Flowering Mar–Apr. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Light dry soils, limestone chip, open rocky areas, among boulders, roadsides, pastures, stony open sandy prairies, dry streamside meadows, calcareous soils, limestone outcroppings, scrub-oak grassland flats |
Elevation | 1500-3500 m (4900-11500 ft) | 150-700 m (500-2300 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
TX |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria recurvata is known from the Edwards Plateau. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 660. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Vesicaria recurvata, Lesquerella recurvata |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Engelmann ex A. Gray) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
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